790:
245:
700:, a nutritious food that the animal is known to place in a shallow cache for later consumption. The habit of caching the seeds is likely to have played an important role in the dispersal of the tree. However, Australian sandalwood was a commercially valuable tree, and was extensively cleared during colonisation. Populations of woylies introduced to an island off the coast of South Australia consume mainly plant material, tubers and roots, seeds and leaves, and beetles, a diet regarded as unusual for the species. Analysis of that colony's diet showed that it included fungal spores, as detected in their
521:
31:
716:. The species was used for meat during the earlier colonial period, although that practice did not persist amongst the colonists because, while it was readily available and easily captured, the skinning of the animal is said to be difficult. At one time, woylies were kept as pets in the same region. Although similar species of marsupials were often regarded as agricultural pests, the woylie did not always acquire this reputation, and was sometimes identified as a non-destructive native animal.
92:
50:
556:. As late as 1910, the population was said to be well known in the Australian south-west, and interviews with older residents helped to establish the time and pattern of decline. The sudden demise of local populations near settlements across the state was noticed by the inhabitants, with most vanishing in the 1930s, while some persisted in a few regions until the 1950s.
1005:. Another 73 animals were translocated from Dryandra National Park in WA in June 2023, when it was reported that the existing bettongs were surviving and breeding. A trapping survey estimated that there are over 200 in Dhilba Guuranda-Innes National Park as a result of the Marna Banggara project as of March 2023.
938:
reserves. In the
Dryandra Woodland, woylie numbers, which had been decimated by hundreds of feral cats, were reported to be recovering after the introduction of a state-sponsored cat baiting and trapping program, supported by the adjacent farming community. It was discovered that most of the feral
815:
A broad foraging area is occupied by individuals, with a larger area for the male, and these will likely overlap with the larger range of other woylies. Within the range, each animal defends a smaller central territory which only overlaps between males and females. Several nests within a range may
896:, which flourished after fox numbers were brought under control. A major program to control feral cats has seen the number of both woylies and numbats begin to increase, although populations remain vastly reduced from even recent decades. In 2011 the species was reported as living mostly in open
412:
marsupial that digs for fungi during the night, usually maintaining a solitary range around a central nest. The length of the head and body combined is 310 to 380 millimetres (12 to 15 in), entirely covered in fur that is a grey-brown over the back, a buff colour across the face, thigh and
876:
infestations in woylie blood. Predation and habitat destruction were also suggested as contributing to the decline of the species. In 2011, the global population was estimated to be less than 5,600 individuals. It was hypothesised in 2013 that disease within populations was making them more
820:
around the nest site for the female and 28 to 43 ha for the male. The central range seemed to extend around the nest for 4 to 6 ha. A subsequent survey calculated an exclusive zone around the nest of 2 to 3 ha for the male, and the foraging area as 27 ha for the male and
811:
The species behaves in a characteristic way when disturbed at its nest, rapidly leaving the site with an explosive noise. The body is arched as the animal hops away, with the head held low and the tail extended, using a bipedal motion in lengthy bounds to evade a potential predator.
816:
provide casual and communal accommodation; only a central nest site is defended to the exclusion of any other. Measurements of the nest territory and foraging ranges of individuals has given variable results. An early study at an open woodland suggested an area of 15 to 28
807:
The woylie builds its dome-shaped nest in a shallow scrape under a bush, and is able to curl its tail to carry bundles of nesting material. The nest, which consists of grass and shredded bark, sticks, leaves and other available material, is well made and discreet.
384:
newspapers, and the spelling 'woylye' was added in the 1920s. The names boodie and boodie rat came to be applied by the rural inhabitants to several species in south-west
Australia, including the woylie, and printed in references to them after 1897. However, the
678:
is consumed by woylies, which avoid the hard outer layer, and has been analysed for its nutritional value. The food is a rich source of lipids and trace elements accumulated by the fungus, and high protein levels that lack the required
430:), although the woylie is distinctly paler at the ventral side, and lacks the blackish colour of the boodie's tail. The ring around the eye of the woylie is pale, and its muzzle is longer and more pointed than the boodie, and less than
417:
is interspersed with silvery hair. The tail is a similar length to the head and body, measuring from 290 to 350 mm (11 to 14 in), and is a rufous brown colour that ends in a blackish tip. The upper-side of the slightly
750:
engineer" by one researcher, moving about six tonnes of soil annually in its quest for food. digging up the soil creating seed beds for new seeds and new plants to grow. This turning over of the soil not only spreads the
743:), which arrived with Europeans, are known to prey on this species, and both have been cited as a major cause of local extinctions. Since the control of the red fox, the cat has become the major predator of the species.
511:
species from their place of birth appears to be by sub-adult males acquiring a new territory. That has been recorded in these species when they reoccupy an area subjected to fire that had killed the previous residents.
864:
in New South Wales. As a result of those efforts, the woylie population rose to sufficient numbers that it was taken off the threatened species list in 1996. The total population of the species rose to 40,000 by 2001.
653:, the bulk of its nutrients are derived from underground fungi, which it digs out with its strong fore-claws. The fungi can only be digested indirectly. They are consumed by bacteria in a portion of its stomach. The
422:
has a ridge of longer fur along its length. The average measurements are 330 mm (13 in) for the heady-body length and 310 mm (12 in) for the tail. The average weight is 1,300 grams (46 oz).
996:
to Dhilba
Guuranda–Innes National Park. Surveillance trapping in 2022 confirmed population survival and breeding. In July 2022, 36 woylies were translocated from WA for the first time, with the permission of the
476:. The population in south-west Australia persisted into the twentieth century, seemingly surviving the mass extinction of similar mammals in the previous decades, and was collected at temperate locations near
785:
before dawn. They can breed all year round if the conditions are favourable. The female is fertile at six months of age and gives birth every 3.5 months. Its lifespan in the wild is about four to six years.
657:
produce the nutrients that are digested in the rest of the animal's stomach and small intestine. When it was widespread and abundant, the woylie probably played an important role in the dispersal of
563:, was in 1912. In some areas, that was recalled as following the earlier disappearance of the boodie. The decline was probably caused by a number of factors, including the impact of introduced
397:, and was sometimes used to distinguish the species from the boodie. Another vernacular term applied to the woylie was "farting rat", inspired by the abrupt noise it emits when disturbed.
988:
of the region. As part of this project, the woylie's reintroduction is being supported by large-scale fencing and the baiting of feral cats and foxes. In August 2021, 40 woylies were
821:
20 ha for the female. An analysis of a radio-tracking survey also indicated a nest range of 2 to 3 ha, yet a smaller foraging range of 7 to 9 ha for the species.
531:
The woylie was originally found in great abundance across southern
Australia, north to about 30°S, in a wide distribution range from the coast in the west and east toward the
312:
in 1837, based on the skin and skull of an adult male obtained by the
Zoological Society of London, and placed with the British Museum of Natural History. The origin of the
868:
However, another sudden decline occurred in late 2001, and in just five years, the woylie population dropped to only 10-30% of its pre-2001 numbers. It was returned to the
927:
931:
1615:
614:
4960:
2113:
833:. By 1992, the woylie was reported from only four small areas in Western Australia. In 1996, it occurred in six sites in Western Australia, including
591:), may have also placed the population under pressure, especially in arid and semi-arid regions, by direct competition or the degradation of habitat.
918:
Despite those declines, woylies continued as small localised populations in predator-free sanctuaries, including a population established in 2010 at
1264:
691:
within the digestion. In other seasons, woylies may adopt a more herbivorous range of foods in addition to that season's hypogeous fungal bodies.
4707:
1535:
1293:
1236:
4919:
4772:
4746:
456:
grasslands. It was abundant in the mid-19th century, inhabiting a range covering around 60% of the
Australian mainland, including all of the
1420:
1340:
1577:
2165:
601:
might also have played a part. The species suffered localised extinctions throughout its range, and was highly endangered by the 1970s.
4970:
841:, and on three islands and two mainland sites in South Australia, following a reintroduction program after foxes had been controlled.
829:
Conservation efforts in the late 20th century concentrated on reintroducing woylies to sites in their former range, after controlling
1830:
872:
as critically endangered. The exact cause of the rapid population crash was uncertain, although researcher Andrew
Thompson found two
2195:
4681:
2083:
1683:
Berry, O.; Angus, J.; Hitchen, Y.; Lawson, J.; Macmahon, B.; Williams, A.A.E.; Thomas, N.D.; Marlow, N.J. (30 April 2015). "Cats (
5000:
4720:
2254:
969:
1753:
1926:
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1208:
489:
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and river itself, and their nests amongst clumps of grass and the hollows of trees, observing a preference for woodlands of
4985:
4932:
4865:
1978:
1630:
2056:
2030:
1952:
1307:
4990:
4980:
2173:
775:, resting during the day and emerging around dusk rather than after sunset, unlike the strictly nocturnal rufous bettong
2138:
2004:
1634:
4995:
3368:
888:
in
Western Australia. The state-sponsored control of foxes offered new hope for the species, but along with the native
4937:
4870:
1818:
1392:
1025:
849:
713:
477:
1190:
4924:
4074:
610:
330:
J.E. Gray, 1837. The nominate subspecies, classified as a modern extinction. This was the south-eastern subspecies.
4777:
4597:
2224:
1900:
1761:
1344:
926:. The reintroduction of control programs for predators saw the species successfully conserved at sites including
904:
838:
4624:
1146:
1345:"Historical perspectives of the ecology of some conspicuous vertebrate species in south-west Western Australia"
1278:
Waterhouse, 1841., G.R. (1841). "The natural history of
Marsupialia or pouched animals". In Jardine, W. (ed.).
993:
939:
cats were living and breeding on surrounding farms, and then entering the woodland to kill the native animals.
853:
3590:
3317:
3233:
3224:
1492:
Short, J. (1998). "The extinction of rat-kangaroos (Marsupialia: Potoroidae) in New South Wales, Australia".
923:
560:
393:) is now recognised as a separate species. The term "kangaroo rat" was also applied from the founding of the
54:
668:, and around 24 fungal taxa are known to be consumed. During the summer months, species of the truffle-like
4101:
3644:
3386:
2227:
1235:
Government of
Western Australia, Department of Biodiversity, Conservation and Attractions (25 April 2022).
4906:
4857:
4852:
4759:
4637:
4632:
4606:
3617:
3608:
2420:
2411:
2247:
1151:
951:
1429:"The status of Australian mammals in 1922 - collections and field notes of museum collector Charles Hoy"
452:
The woylie is known to have once inhabited a wide range of habitats, including low arid scrub or desert
4200:
4173:
4110:
3957:
3572:
3159:
3150:
1844:
Williams, M.R.; Wayne, J.C.; Wilson, I.; Vellios, C.V.; Ward, C.G.; Maxwell, M.A.; Wayne, A.F. (2013).
1287:
845:
789:
493:
91:
4911:
4191:
4155:
3948:
3395:
2790:
2772:
543:
380:. The spelling woylie, and its variants, were used for the species in reports and advertisements in
288:
and carries its young in a pouch. Once widespread, the woylie mostly died out from habitat loss and
4209:
4146:
4047:
3581:
3344:
3299:
2901:
2799:
2707:
1536:"Numbats and woylies flourish at Dryandra after feral cats pushed WA icon towards 'extinction pit'"
799:
1073:
413:
flank, and blending to the pale cream colour beneath. The greyish brown of the upper-parts of the
4092:
3796:
3713:
3671:
3404:
3308:
3290:
3281:
3209:
3200:
2808:
2671:
893:
580:
353:
292:. It is currently restricted to two small areas in Western Australia. There were two subspecies:
244:
4803:
496:
regions of Western Australia in the 1930s coincided with the earliest records of the introduced
4712:
4559:
4164:
4137:
3755:
3653:
3599:
3326:
3257:
3248:
3191:
3053:
3044:
2919:
2629:
2611:
2602:
2240:
1133:
Magazine of Natural History and Journal of Zoology, Botany, Mineralogy, Geology and Meteorology
943:
830:
576:
1282:. Vol. 11 (1 ed.). Edinburgh & London: W.H. Lizars & H.G. Bohn. p. 185.
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4611:
4292:
3889:
3871:
3838:
3829:
3465:
3456:
3377:
3335:
3086:
2877:
2731:
2680:
2566:
1888:
1826:
989:
777:
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and seeds of other native plants, but provides good beds for these the seeds to grow, so the
196:
683:
and other amino acids, may be compensated by fermentation processes on the available aminos
520:
4790:
4668:
4489:
4319:
4310:
4274:
4182:
3680:
2928:
2910:
2698:
2515:
2200:
1501:
912:
532:
431:
289:
4738:
2166:"Brush-tailed bettongs flown from Western Australia get a new home on the Yorke Peninsula"
1198:
8:
4507:
4471:
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4371:
4265:
4128:
3981:
3805:
3635:
3540:
3531:
3011:
2832:
2823:
834:
609:
As of 2021, there are only two indigenous populations of woylies, one of which is in the
469:
457:
39:
1505:
4965:
4526:
4517:
4480:
4301:
3916:
3441:
3432:
3002:
2781:
2506:
1800:
1720:
985:
962:
947:
919:
727:), a large raptor thought to have been a significant influence on their mortality. The
720:
547:
86:
1513:
4785:
4645:
3931:
2647:
1871:
1765:
1712:
1450:
1388:
1258:
1214:
1204:
1021:
1002:
935:
881:
696:
552:
539:
461:
394:
381:
336:(Waterhouse, 1841). This is the south-western subspecies. A description published as
1724:
1050:
942:
The species has become established at a large fenced reserve at Western Australia's
4650:
4439:
3907:
2755:
2593:
2584:
2452:
1861:
1704:
1509:
1440:
1068:
861:
639:
diet and will dig for a wide variety of their fruiting bodies. Although it may eat
546:
visited during its founding years. Gilbert noted woylies on the tidal flats of the
481:
365:
345:
276:
native to forests and shrubland of Australia. A member of the rat-kangaroo family (
212:
664:
The austral summer and autumn seasons provide woylies with the fruiting bodies of
30:
4897:
4843:
4412:
4283:
3999:
3764:
3731:
3704:
3497:
3140:
3122:
3113:
3077:
3020:
2575:
2548:
2323:
1376:
981:
973:
665:
584:
473:
465:
453:
419:
317:
4816:
4751:
2084:"Controversial wire fence splits peninsula to keep native animals in, pests out"
504:), which was first seen at Perth in 1927, and in the south not long thereafter.
3990:
3880:
3722:
3272:
3174:
3029:
2740:
2689:
2232:
1124:
977:
958:
704:, although that is likely to have been an occupant of the guts of the beetles.
617:, and it has been found that their numbers have risen in the years up to 2021.
485:
1866:
1845:
4954:
4733:
4582:
4343:
4334:
3898:
3862:
3555:
3359:
2620:
2557:
2467:
2387:
2378:
2333:
2311:
2264:
2114:"Endangered bettongs complete 2,000km journey to new home in South Australia"
2031:"Rewilding project to create 'great southern ark' over former mine, farmland"
1875:
1716:
1454:
1194:
1059:
869:
794:
649:
153:
59:
1578:"Meet the woylie, an eco-engineer bringing life back to degraded ecosystems"
1218:
694:
Woylies have been observed eating the large seeds of Australian sandalwood,
4023:
4008:
3853:
3779:
3509:
3104:
2993:
2965:
2956:
2946:
2850:
2722:
2638:
1380:
1186:
1095:
658:
525:
4694:
4591:
3746:
3422:
3095:
2892:
2662:
2539:
2489:
2402:
1424:
908:
897:
670:
636:
598:
572:
568:
4660:
1445:
1428:
4686:
4361:
3972:
2983:
880:
By 2010, one of the remaining thriving habitats for the woylie was the
688:
349:
309:
277:
163:
4699:
4248:
3695:
2530:
2295:
2196:"Yorke Peninsula's brush-tailed bettong population continues to grow"
857:
772:
747:
736:
631:
508:
409:
285:
143:
103:
4553:
2082:
Corvo, Shannon; Radford, Luke; Lysaght, Gary-Jon (7 November 2019).
1708:
968:
It is a key species in the faunal reconstruction project within the
4891:
4837:
4576:
4065:
3820:
3068:
2868:
2283:
873:
760:
684:
654:
313:
123:
957:
As of 2020, plans were in place to reintroduce the species to the
4454:
4395:
3662:
3626:
3517:
1387:(3rd ed.). Melbourne: Oxford University Press. p. 106.
1200:
Mammal Species of the World: A Taxonomic and Geographic Reference
998:
817:
728:
626:
564:
497:
173:
438:), with which it once shared an overlapping distribution range.
4673:
4619:
4421:
4233:
4224:
4056:
2475:
2360:
2289:
2277:
1234:
1044:
1042:
1040:
900:
889:
680:
644:
414:
386:
281:
273:
133:
113:
1687:) are more abundant and are the dominant predator of woylies (
1147:"Subspecies Bettongia penicillata penicillata J.E. Gray, 1837"
4811:
2342:
885:
756:
752:
701:
674:
form the major part of the diet. The core of the truffles of
640:
594:
By the 1920s, the woylie was extinct over much of its range.
368:. The regional variants amongst Nyungar peoples are noted as
76:
70:
1203:(3rd ed.). Johns Hopkins University Press. p. 57.
1037:
340:, a species now recognised as the only extant subspecies of
1652:
1048:
782:
280:), it moves by hopping and is active at night, digging for
1927:"Woylies make a historic return to the Northern Territory"
1843:
1780:
1554:
1473:
1461:
1633:- Department of Land Resource Management. Archived from
976:, formerly named the Great Southern Ark (2019), renamed
615:
Department of Biodiversity, Conservation and Attractions
2450:
1979:"Benchmarking AWC's progress at two NSW National Parks"
1682:
1664:
1596:
583:
has undoubtedly been crucial. The introduction of the
1731:
844:
Stable populations were established in places such as
252:
Historic woylie range in yellow, current range in red
1953:"Woylies reintroduced to Mallee Cliffs National Park"
1401:
1846:"Sudden and rapid decline of the abundant marsupial
1167:
946:. It was reintroduced to large fenced landscapes at
2081:
2057:"Great Southern Ark project renamed Marna Banggara"
1764:- Department of Parks and Wildlife. Archived from
714:first European settlers of rural Western Australia
316:has not been determined, but it is presumed to be
1277:
4952:
2262:
1571:
1569:
1371:
1369:
1367:
1365:
1016:Claridge, A.W.; Seebeck, J.H.; Rose, R. (2007).
950:in the Northern Territory in August 2021 and to
2107:
2105:
1375:
1308:"Aboriginal Words in the English Language: L-Z"
1018:Bettongs, potoroos, and the musky rat-kangaroo
4515:
2248:
1798:
1566:
1362:
1335:
1333:
1331:
1329:
1327:
1325:
1271:
1074:10.2305/IUCN.UK.2016-2.RLTS.T2785A21961347.en
538:The species was observed at all parts of the
426:This species resembles the burrowing boodie (
3495:
2331:
2102:
1901:"Endangered Woylies increasing at Mt Gibson"
1837:
1627:Threatened species of the Northern Territory
1263:: CS1 maint: multiple names: authors list (
613:of WA. This group is being monitored by the
4961:IUCN Red List critically endangered species
1748:
1746:
1419:
1292:: CS1 maint: numeric names: authors list (
746:Woylies are natural excavators, dubbed an "
719:Native predators of the woylie include the
567:animals, accompanied by land clearance for
441:
360:The common name, "woylie", is derived from
2309:
2255:
2241:
1529:
1527:
1525:
1523:
1413:
1322:
1118:
1116:
781:, although woylies always return to their
243:
48:
29:
1865:
1676:
1485:
1444:
1385:A field guide to the mammals of Australia
1072:
348:, who presented a manuscript prepared by
1786:
1743:
1737:
1670:
1658:
1602:
1575:
1560:
1479:
1467:
1407:
1230:
1228:
1173:
1015:
788:
519:
2954:
2193:
2163:
2111:
2028:
1792:
1533:
1520:
1113:
907:eucalypt assemblages, with a dense low
4953:
1584:. Australian Broadcasting Corporation)
1339:
1185:
1049:Woinarski, J.; Burbidge, A.A. (2016).
515:
488:in 1920. The last collections made at
4558:
4557:
3494:
2449:
2308:
2236:
2090:. Australian Broadcasting Corporation
2037:. Australian Broadcasting Corporation
1983:AWC - Australian Wildlife Conservancy
1957:AWC - Australian Wildlife Conservancy
1931:AWC - Australian Wildlife Conservancy
1905:AWC - Australian Wildlife Conservancy
1811:
1608:
1542:. Australian Broadcasting Corporation
1534:Kennedy, Elicia (22 September 2019).
1491:
1225:
2164:Cominos, Christian (1 August 2022).
2139:"Bettongs doing well on SA mainland"
1122:
1020:. Collingwood, Victoria: CSIRO Pub.
472:, and across the central portion of
272:) is a small, critically endangered
2194:Cominos, Christian (20 June 2023).
2174:Australian Broadcasting Corporation
1179:
1060:IUCN Red List of Threatened Species
970:Dhilba Guuranda–Innes National Park
323:The two subspecies recognised are:
13:
4075:Eastern short-eared rock-wallaby (
1310:. One Big Garden. 13 December 2011
1280:The Naturalist's Library. Mammalia
1139:
635:species, the woylie has a largely
559:The last sighting of a woylie, at
290:introduced predators such as foxes
14:
5012:
4971:Mammals of the Northern Territory
4858:Bettongia_penicillata_penicillata
4829:Bettongia penicillata penicillata
2218:
2029:Kilvert, Nick (31 January 2019).
1001:traditional custodians, to boost
447:
328:Bettongia penicillata penicillata
308:A species was first described by
296:in the west, and the now-extinct
1799:Leonie Harris (7 October 2008).
1754:"Woylie (Bettongia penicillata)"
90:
3318:Herbert River ringtail possum (
3234:Rock-haunting ringtail possum (
2225:Australian Wildlife Conservancy
2187:
2157:
2131:
2112:Johnson, Sarah (28 July 2022).
2075:
2049:
2022:
2005:"Woylie (Brush-tailed Bettong)"
1997:
1971:
1945:
1919:
1893:
1882:
1801:"Woylie marsupial under threat"
1762:Government of Western Australia
1576:Stephens, Kate (4 April 2021).
905:Mallee Woodlands and Shrublands
839:Australian Wildlife Conservancy
824:
712:Woylies were well known to the
5001:Taxa named by John Edward Gray
3645:Golden-mantled tree-kangaroo (
3387:Plush-coated ringtail possum (
3369:D'Albertis's ringtail possum (
1300:
1088:
980:in June 2020 in honour of the
400:
1:
4912:Bettongia_penicillata_ogilbyi
4883:Bettongia penicillata ogilbyi
2421:Northern hairy-nosed wombat (
2412:Southern hairy-nosed wombat (
1831:Department of the Environment
1821:Bettongia penicillata ogilbyi
1758:Shark Bay World Heritage Area
1697:Australian Journal of Zoology
1631:Northern Territory Government
1616:"Brush-tailed Bettong Woylie
1514:10.1016/s0006-3207(98)00026-3
1008:
561:Bridgetown, Western Australia
334:Bettongia penicillata ogilbyi
4976:Mammals of Western Australia
4201:Purple-necked rock-wallaby (
4174:Black-flanked rock-wallaby (
4111:Yellow-footed rock-wallaby (
3958:Northern nail-tail wallaby (
3591:Goodfellow's tree-kangaroo (
3160:Lemur-like ringtail possum (
1352:Conservation Science W. Aust
924:Western Australian wheatbelt
892:, numbers were decimated by
766:
352:. The type was collected at
7:
4986:Mammals of Victoria (state)
4192:Brush-tailed rock-wallaby (
4102:Rothschild's rock-wallaby (
3949:Bridled nail-tail wallaby (
3396:Reclusive ringtail possum (
2791:Mountain brushtail possum (
2773:Northern brushtail possum (
1152:Australian Faunal Directory
952:Mallee Cliffs National Park
468:, the north-west corner of
303:
298:B. p. penicillata
10:
5017:
4991:Endemic fauna of Australia
4981:Mammals of South Australia
4210:Mount Claro rock-wallaby (
4048:Short-eared rock-wallaby (
3618:Matschie's tree-kangaroo (
3609:Lumholtz's tree-kangaroo (
3345:Vogelkop ringtail possum (
3300:Cinereus ringtail possum (
3210:Southern greater glider (
2902:Long-tailed pygmy possum (
2800:Coppery brushtail possum (
2708:Blue-eyed spotted cuscus (
954:in NSW in September 2021.
860:in Western Australia, and
707:
661:within desert ecosystems.
604:
445:
235:B. penicillata penicillata
4996:Mammals described in 1837
4881:
4827:
4566:
4506:
4452:
4393:
4369:
4360:
4332:
4246:
4222:
4093:Proserpine rock-wallaby (
4021:
3970:
3929:
3851:
3818:
3797:Spectacled hare-wallaby (
3777:
3744:
3714:White-striped dorcopsis (
3693:
3573:Bennett's tree-kangaroo (
3553:
3529:
3508:
3504:
3490:
3454:
3430:
3421:
3405:Coppery ringtail possum (
3357:
3309:Painted ringtail possum (
3291:Weyland ringtail possum (
3282:Lowland ringtail possum (
3270:
3246:
3222:
3201:Northern greater glider (
3172:
3148:
3139:
3066:
3042:
2991:
2982:
2945:
2890:
2866:
2849:
2821:
2809:Common brushtail possum (
2753:
2720:
2672:Admiralty Island cuscus (
2660:
2528:
2487:
2466:
2462:
2445:
2400:
2376:
2359:
2322:
2318:
2304:
2272:
1867:10.1017/S0030605313000677
877:vulnerable to predation.
251:
242:
225:
220:
202:
195:
87:Scientific classification
85:
68:
46:
37:
28:
23:
4165:Unadorned rock-wallaby (
4156:Herbert's rock-wallaby (
4138:Cape York rock-wallaby (
4123:P. lateralis/penicillata
3654:Lowlands tree-kangaroo (
3600:Grizzled tree-kangaroo (
3327:Masked ringtail possum (
3258:Common ringtail possum (
3192:Central greater glider (
2920:Tasmanian pygmy possum (
2612:Northern common cuscus (
2603:Southern common cuscus (
1135:. 1 New Series (2): 584.
647:, and resin exuded from
442:Distribution and habitat
4147:Godman's rock-wallaby (
3890:Western brush wallaby (
3872:Black-striped wallaby (
3839:Eastern grey kangaroo (
3830:Western grey kangaroo (
3582:Doria's tree-kangaroo (
3466:Feather-tailed possum (
3378:Green ringtail possum (
3336:Pygmy ringtail possum (
3087:Yellow-bellied glider (
2878:Mountain pygmy possum (
2732:Sulawesi dwarf cuscus (
2681:Common spotted cuscus (
2567:Eastern common cuscus (
1691:) after sustained fox (
1494:Biological Conservation
1197:; Reeder, D. M (eds.).
620:
354:York, Western Australia
294:B. p. ogilbyi
4320:Red-necked pademelon (
4311:Red-legged pademelon (
4183:Mareeba rock-wallaby (
3681:Ursine tree-kangaroo (
3672:Seri's tree-kangaroo (
2929:Eastern pygmy possum (
2911:Western pygmy possum (
2699:Black-spotted cuscus (
2516:Sulawesi bear cuscus (
1237:"Woylie Fauna Profile"
944:Mount Gibson Sanctuary
804:
542:when the field worker
528:
507:The dispersal of many
344:. The cited author is
229:B. penicillata ogilbyi
4799:Paleobiology Database
4625:bettongia-penicillata
4612:Bettongia_penicillata
4598:Bettongia penicillata
4568:Bettongia penicillata
4472:Long-footed potoroo (
4381:Rufous rat-kangaroo (
4266:Tasmanian pademelon (
4129:Allied rock-wallaby (
3982:Antilopine kangaroo (
3939:(nail-tail wallabies)
3806:Rufous hare-wallaby (
3756:Macleay's dorcopsis (
3541:Banded hare-wallaby (
3054:Leadbeater's possum (
3012:Long-fingered triok (
2833:Scaly-tailed possum (
2630:Rothschild's cuscus (
1848:Bettongia penicillata
1827:Australian Government
1689:Bettongia penicillata
1618:Bettongia penicillata
1191:"Order Diprotodontia"
1155:. biodiversity.org.au
1127:Bettongia penicillata
1053:Bettongia penicillata
972:on South Australia's
792:
778:Aepyprymnus rufescens
589:Oryctolagus cuniculus
523:
406:Bettongia penicillata
284:to eat. It is also a
269:Bettongia penicillata
206:Bettongia penicillata
55:Critically Endangered
4527:Musky rat-kangaroo (
4481:Long-nosed potoroo (
4302:Mountain pademelon (
4293:Calaby's pademelon (
3917:Red-necked wallaby (
3442:Feathertail glider (
3003:Great-tailed triok (
2782:Short-eared possum (
2507:Talaud bear cuscus (
2201:ABC News (Australia)
1433:Australian Zoologist
1096:"Appendices | CITES"
1067:: e.T2785A21961347.
856:in South Australia,
800:Mammals of Australia
771:Woylies are largely
577:introduced red foxes
533:Great Dividing Range
338:Hypsiprymnus ogilbyi
264:brush-tailed bettong
16:Species of marsupial
4508:Hypsiprymnodontidae
4490:Gilbert's potoroo (
4275:Brown's pademelon (
2763:(brushtail possums)
2569:P. intercastellanus
1661:, pp. 116–117.
1506:1998BCons..86..365S
1446:10.7882/az.2001.002
1123:Gray, J.E. (1837).
835:Karakamia Sanctuary
611:Upper Warren region
516:Population collapse
448:§ Conservation
342:B. penicillata
188:B. penicillata
40:Conservation status
4440:Northern bettong (
3908:Whiptail wallaby (
2594:Telefomin cuscus (
2585:Blue-eyed cuscus (
2063:. 27 November 2020
1768:on 2 December 2013
1244:www.dpaw.wa.gov.au
986:traditional owners
963:Dirk Hartog Island
920:Wadderin Sanctuary
805:
721:wedge-tailed eagle
548:Swan Coastal Plain
529:
436:Potorous gilbertii
382:Western Australian
300:in the southeast.
4948:
4947:
4786:Open Tree of Life
4560:Taxon identifiers
4551:
4550:
4547:
4546:
4543:
4542:
4539:
4538:
4502:
4501:
4465:
4413:Eastern bettong (
4406:
4356:
4355:
4284:Dusky pademelon (
4259:
4203:P. purpureicollis
4034:
4000:Common wallaroo (
3942:
3799:L. conspicillatus
3790:
3765:Small dorcopsis (
3732:Brown dorcopsis (
3705:Black dorcopsis (
3566:
3523:
3486:
3485:
3482:
3481:
3478:
3477:
3417:
3416:
3185:
3182:(greater gliders)
3135:
3134:
3123:Squirrel glider (
3114:Mahogany glider (
3078:Northern glider (
2978:
2977:
2941:
2940:
2860:
2845:
2844:
2766:
2576:Woodlark cuscus (
2549:Mountain cuscus (
2500:
2481:
2456:
2453:Phalangeriformes
2441:
2440:
2437:
2436:
2433:
2432:
2370:
2355:
2354:
2228:woylie fact sheet
1959:. 9 February 2022
1789:, pp. 76–77.
1563:, pp. 83–84.
1482:, pp. 14–15.
1470:, pp. 82–83.
1288:cite encyclopedia
1210:978-0-8018-8221-0
1003:genetic diversity
936:Dryandra Woodland
882:Dryandra Woodland
763:is kept healthy.
697:Santalum spicatum
553:Eucalyptus wandoo
540:Swan River Colony
490:King George Sound
462:Western Australia
432:Gilbert's potoroo
428:Bettongia lesueur
395:Swan River Colony
391:Bettongia lesueur
256:
255:
80:
63:
5008:
4941:
4940:
4928:
4927:
4915:
4914:
4902:
4901:
4900:
4874:
4873:
4861:
4860:
4848:
4847:
4846:
4820:
4819:
4807:
4806:
4794:
4793:
4781:
4780:
4768:
4767:
4755:
4754:
4742:
4741:
4729:
4728:
4716:
4715:
4703:
4702:
4690:
4689:
4677:
4676:
4664:
4663:
4654:
4653:
4641:
4640:
4628:
4627:
4615:
4614:
4602:
4601:
4600:
4587:
4586:
4585:
4555:
4554:
4513:
4512:
4464:
4463:
4459:
4405:
4404:
4400:
4367:
4366:
4258:
4257:
4253:
4033:
4032:
4031:(rock-wallabies)
4028:
3991:Black wallaroo (
3941:
3940:
3936:
3881:Tammar wallaby (
3789:
3788:
3787:(hare-wallabies)
3784:
3723:Gray dorcopsis (
3565:
3564:
3563:(tree-kangaroos)
3560:
3527:
3526:
3522:
3521:
3513:
3506:
3505:
3492:
3491:
3428:
3427:
3184:
3183:
3179:
3146:
3145:
3030:Striped possum (
2989:
2988:
2952:
2951:
2864:
2863:
2859:
2858:
2854:
2835:W. squamicaudata
2765:
2764:
2760:
2741:Banggai cuscus (
2690:Waigeou cuscus (
2648:Stein's cuscus (
2499:
2498:
2494:
2485:
2484:
2480:
2479:
2471:
2464:
2463:
2454:
2447:
2446:
2374:
2373:
2369:
2368:
2364:
2329:
2328:
2320:
2319:
2306:
2305:
2257:
2250:
2243:
2234:
2233:
2213:
2212:
2210:
2208:
2191:
2185:
2184:
2182:
2180:
2161:
2155:
2154:
2152:
2150:
2135:
2129:
2128:
2126:
2124:
2109:
2100:
2099:
2097:
2095:
2079:
2073:
2072:
2070:
2068:
2053:
2047:
2046:
2044:
2042:
2026:
2020:
2019:
2017:
2015:
2001:
1995:
1994:
1992:
1990:
1975:
1969:
1968:
1966:
1964:
1949:
1943:
1942:
1940:
1938:
1923:
1917:
1916:
1914:
1912:
1897:
1891:
1886:
1880:
1879:
1869:
1841:
1835:
1834:
1815:
1809:
1808:
1796:
1790:
1784:
1778:
1777:
1775:
1773:
1750:
1741:
1735:
1729:
1728:
1680:
1674:
1668:
1662:
1656:
1650:
1649:
1647:
1645:
1639:
1624:
1612:
1606:
1600:
1594:
1593:
1591:
1589:
1573:
1564:
1558:
1552:
1551:
1549:
1547:
1531:
1518:
1517:
1489:
1483:
1477:
1471:
1465:
1459:
1458:
1448:
1417:
1411:
1405:
1399:
1398:
1373:
1360:
1359:
1349:
1337:
1320:
1319:
1317:
1315:
1304:
1298:
1297:
1291:
1283:
1275:
1269:
1268:
1262:
1254:
1252:
1250:
1241:
1232:
1223:
1222:
1183:
1177:
1171:
1165:
1164:
1162:
1160:
1143:
1137:
1136:
1120:
1111:
1110:
1108:
1106:
1092:
1086:
1085:
1083:
1081:
1076:
1046:
1031:
862:Scotia Sanctuary
793:Illustration by
484:in 1909, and by
482:G. C. Shortridge
408:is a species of
366:Nyungar language
346:G. R. Waterhouse
247:
208:
95:
94:
74:
57:
52:
51:
33:
21:
20:
5016:
5015:
5011:
5010:
5009:
5007:
5006:
5005:
4951:
4950:
4949:
4944:
4936:
4931:
4923:
4918:
4910:
4905:
4896:
4895:
4890:
4877:
4869:
4864:
4856:
4851:
4842:
4841:
4836:
4823:
4815:
4810:
4802:
4797:
4789:
4784:
4776:
4771:
4763:
4758:
4750:
4745:
4737:
4732:
4724:
4719:
4711:
4706:
4698:
4693:
4685:
4680:
4672:
4667:
4659:
4657:
4649:
4644:
4636:
4631:
4623:
4618:
4610:
4605:
4596:
4595:
4590:
4581:
4580:
4575:
4562:
4552:
4535:
4498:
4461:
4460:
4458:
4448:
4402:
4401:
4399:
4389:
4352:
4344:Swamp wallaby (
4328:
4268:T. billardierii
4255:
4254:
4252:
4242:
4218:
4030:
4029:
4027:
4017:
3966:
3938:
3937:
3935:
3925:
3899:Parma wallaby (
3863:Agile wallaby (
3847:
3814:
3786:
3785:
3783:
3773:
3740:
3689:
3647:D. pulcherrimus
3575:D. bennettianus
3562:
3561:
3559:
3549:
3515:
3514:
3512:
3500:
3498:Macropodiformes
3474:
3450:
3413:
3353:
3320:P. herbertensis
3266:
3242:
3218:
3181:
3180:
3178:
3168:
3141:Pseudocheiridae
3131:
3125:P. norfolcensis
3062:
3038:
2974:
2937:
2886:
2857:(pygmy possums)
2856:
2855:
2853:
2841:
2817:
2762:
2761:
2759:
2749:
2716:
2656:
2621:Ornate cuscus (
2558:Ground cuscus (
2524:
2497:(bear cuscuses)
2496:
2495:
2493:
2473:
2472:
2470:
2458:
2429:
2396:
2388:Common wombat (
2366:
2365:
2363:
2351:
2324:Phascolarctidae
2314:
2300:
2268:
2261:
2221:
2216:
2206:
2204:
2192:
2188:
2178:
2176:
2162:
2158:
2148:
2146:
2137:
2136:
2132:
2122:
2120:
2110:
2103:
2093:
2091:
2080:
2076:
2066:
2064:
2055:
2054:
2050:
2040:
2038:
2027:
2023:
2013:
2011:
2003:
2002:
1998:
1988:
1986:
1977:
1976:
1972:
1962:
1960:
1951:
1950:
1946:
1936:
1934:
1933:. 5 August 2021
1925:
1924:
1920:
1910:
1908:
1899:
1898:
1894:
1887:
1883:
1842:
1838:
1817:
1816:
1812:
1797:
1793:
1785:
1781:
1771:
1769:
1752:
1751:
1744:
1736:
1732:
1709:10.1071/ZO14024
1681:
1677:
1669:
1665:
1657:
1653:
1643:
1641:
1637:
1622:
1614:
1613:
1609:
1601:
1597:
1587:
1585:
1574:
1567:
1559:
1555:
1545:
1543:
1532:
1521:
1490:
1486:
1478:
1474:
1466:
1462:
1418:
1414:
1406:
1402:
1395:
1377:Menkhorst, P.W.
1374:
1363:
1347:
1338:
1323:
1313:
1311:
1306:
1305:
1301:
1285:
1284:
1276:
1272:
1256:
1255:
1248:
1246:
1239:
1233:
1226:
1211:
1184:
1180:
1172:
1168:
1158:
1156:
1145:
1144:
1140:
1121:
1114:
1104:
1102:
1094:
1093:
1089:
1079:
1077:
1047:
1038:
1034:
1028:
1011:
982:Narungga people
974:Yorke Peninsula
961:in NSW, and to
922:in the central
913:tussock grasses
850:St Peter Island
827:
769:
710:
666:hypogeous fungi
623:
607:
585:European rabbit
575:. Predation by
518:
474:New South Wales
466:South Australia
450:
444:
420:prehensile tail
403:
318:New South Wales
306:
216:
210:
204:
191:
89:
81:
64:
53:
49:
42:
17:
12:
11:
5:
5014:
5004:
5003:
4998:
4993:
4988:
4983:
4978:
4973:
4968:
4963:
4946:
4945:
4943:
4942:
4929:
4916:
4903:
4887:
4885:
4879:
4878:
4876:
4875:
4862:
4849:
4833:
4831:
4825:
4824:
4822:
4821:
4808:
4795:
4782:
4769:
4756:
4743:
4730:
4717:
4704:
4691:
4678:
4665:
4655:
4642:
4629:
4616:
4603:
4588:
4572:
4570:
4564:
4563:
4549:
4548:
4545:
4544:
4541:
4540:
4537:
4536:
4534:
4533:
4523:
4521:
4518:Hypsiprymnodon
4510:
4504:
4503:
4500:
4499:
4497:
4496:
4487:
4483:P. tridactylus
4478:
4468:
4466:
4450:
4449:
4447:
4446:
4437:
4433:B. penicillata
4428:
4419:
4409:
4407:
4391:
4390:
4388:
4387:
4377:
4375:
4364:
4358:
4357:
4354:
4353:
4351:
4350:
4340:
4338:
4330:
4329:
4327:
4326:
4317:
4308:
4299:
4290:
4281:
4272:
4262:
4260:
4244:
4243:
4241:
4240:
4230:
4228:
4220:
4219:
4217:
4216:
4207:
4198:
4194:P. penicillata
4189:
4180:
4171:
4162:
4153:
4144:
4135:
4118:
4117:
4108:
4104:P. rothschildi
4099:
4082:
4081:
4072:
4063:
4054:
4037:
4035:
4019:
4018:
4016:
4015:
4009:Red kangaroo (
4006:
3997:
3988:
3984:O. antilopinus
3978:
3976:
3968:
3967:
3965:
3964:
3955:
3945:
3943:
3927:
3926:
3924:
3923:
3919:N. rufogriseus
3914:
3905:
3896:
3887:
3878:
3869:
3859:
3857:
3849:
3848:
3846:
3845:
3836:
3832:M. fuliginosus
3826:
3824:
3816:
3815:
3813:
3812:
3803:
3793:
3791:
3775:
3774:
3772:
3771:
3762:
3752:
3750:
3742:
3741:
3739:
3738:
3729:
3720:
3711:
3701:
3699:
3691:
3690:
3688:
3687:
3678:
3669:
3660:
3651:
3642:
3633:
3624:
3615:
3606:
3597:
3593:D. goodfellowi
3588:
3579:
3569:
3567:
3551:
3550:
3548:
3547:
3537:
3535:
3524:
3502:
3501:
3488:
3487:
3484:
3483:
3480:
3479:
3476:
3475:
3473:
3472:
3462:
3460:
3452:
3451:
3449:
3448:
3438:
3436:
3425:
3419:
3418:
3415:
3414:
3412:
3411:
3402:
3393:
3384:
3375:
3365:
3363:
3355:
3354:
3352:
3351:
3342:
3333:
3324:
3315:
3306:
3297:
3288:
3278:
3276:
3273:Pseudochirulus
3268:
3267:
3265:
3264:
3254:
3252:
3244:
3243:
3241:
3240:
3230:
3228:
3220:
3219:
3217:
3216:
3207:
3198:
3188:
3186:
3170:
3169:
3167:
3166:
3156:
3154:
3143:
3137:
3136:
3133:
3132:
3130:
3129:
3120:
3111:
3105:Sugar glider (
3102:
3093:
3084:
3074:
3072:
3064:
3063:
3061:
3060:
3056:G. leadbeateri
3050:
3048:
3040:
3039:
3037:
3036:
3027:
3021:Tate's triok (
3018:
3009:
2999:
2997:
2986:
2980:
2979:
2976:
2975:
2973:
2972:
2966:Honey possum (
2962:
2960:
2949:
2943:
2942:
2939:
2938:
2936:
2935:
2926:
2917:
2908:
2898:
2896:
2888:
2887:
2885:
2884:
2874:
2872:
2861:
2847:
2846:
2843:
2842:
2840:
2839:
2829:
2827:
2819:
2818:
2816:
2815:
2806:
2797:
2793:T. cunninghami
2788:
2779:
2775:T. arnhemensis
2769:
2767:
2751:
2750:
2748:
2747:
2738:
2728:
2726:
2718:
2717:
2715:
2714:
2705:
2696:
2687:
2678:
2668:
2666:
2658:
2657:
2655:
2654:
2645:
2639:Silky cuscus (
2636:
2632:P. rothschildi
2627:
2618:
2609:
2600:
2591:
2582:
2573:
2564:
2555:
2546:
2536:
2534:
2526:
2525:
2523:
2522:
2513:
2503:
2501:
2482:
2460:
2459:
2443:
2442:
2439:
2438:
2435:
2434:
2431:
2430:
2428:
2427:
2418:
2408:
2406:
2398:
2397:
2395:
2394:
2384:
2382:
2371:
2357:
2356:
2353:
2352:
2350:
2349:
2339:
2337:
2326:
2316:
2315:
2302:
2301:
2299:
2298:
2292:
2286:
2280:
2273:
2270:
2269:
2260:
2259:
2252:
2245:
2237:
2231:
2230:
2220:
2219:External links
2217:
2215:
2214:
2186:
2156:
2130:
2118:Marna Banggara
2101:
2074:
2048:
2021:
1996:
1970:
1944:
1918:
1892:
1881:
1860:(1): 175–185.
1836:
1810:
1791:
1779:
1742:
1730:
1675:
1673:, p. 108.
1663:
1651:
1640:on 19 May 2014
1607:
1605:, p. 104.
1595:
1565:
1553:
1519:
1500:(3): 365–377.
1484:
1472:
1460:
1439:(4): 533–562.
1412:
1400:
1393:
1361:
1321:
1299:
1270:
1224:
1209:
1178:
1166:
1138:
1112:
1087:
1035:
1033:
1032:
1026:
1012:
1010:
1007:
984:, who are the
978:Marna Banggara
959:Pilliga Forest
826:
823:
768:
765:
709:
706:
622:
619:
606:
603:
517:
514:
486:Charles M. Hoy
478:Margaret River
443:
440:
402:
399:
358:
357:
331:
305:
302:
254:
253:
249:
248:
240:
239:
238:
237:
231:
223:
222:
218:
217:
211:
200:
199:
193:
192:
185:
183:
179:
178:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
121:
117:
116:
111:
107:
106:
101:
97:
96:
83:
82:
69:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
5013:
5002:
4999:
4997:
4994:
4992:
4989:
4987:
4984:
4982:
4979:
4977:
4974:
4972:
4969:
4967:
4964:
4962:
4959:
4958:
4956:
4939:
4934:
4930:
4926:
4921:
4917:
4913:
4908:
4904:
4899:
4893:
4889:
4888:
4886:
4884:
4880:
4872:
4867:
4863:
4859:
4854:
4850:
4845:
4839:
4835:
4834:
4832:
4830:
4826:
4818:
4813:
4809:
4805:
4800:
4796:
4792:
4787:
4783:
4779:
4774:
4770:
4766:
4761:
4757:
4753:
4748:
4744:
4740:
4735:
4731:
4727:
4722:
4718:
4714:
4709:
4705:
4701:
4696:
4692:
4688:
4683:
4679:
4675:
4670:
4666:
4662:
4656:
4652:
4647:
4643:
4639:
4634:
4630:
4626:
4621:
4617:
4613:
4608:
4604:
4599:
4593:
4589:
4584:
4578:
4574:
4573:
4571:
4569:
4565:
4561:
4556:
4532:
4530:
4525:
4524:
4522:
4520:
4519:
4514:
4511:
4509:
4505:
4495:
4493:
4488:
4486:
4484:
4479:
4477:
4475:
4470:
4469:
4467:
4457:
4456:
4451:
4445:
4443:
4438:
4436:
4434:
4429:
4427:
4425:
4420:
4418:
4416:
4411:
4410:
4408:
4398:
4397:
4392:
4386:
4384:
4379:
4378:
4376:
4374:
4373:
4368:
4365:
4363:
4359:
4349:
4347:
4342:
4341:
4339:
4337:
4336:
4331:
4325:
4323:
4318:
4316:
4314:
4313:T. stigmatica
4309:
4307:
4305:
4300:
4298:
4296:
4291:
4289:
4287:
4282:
4280:
4278:
4273:
4271:
4269:
4264:
4263:
4261:
4251:
4250:
4245:
4239:
4237:
4236:S. brachyurus
4232:
4231:
4229:
4227:
4226:
4221:
4215:
4213:
4208:
4206:
4204:
4199:
4197:
4195:
4190:
4188:
4186:
4181:
4179:
4177:
4172:
4170:
4168:
4163:
4161:
4159:
4154:
4152:
4150:
4145:
4143:
4141:
4136:
4134:
4132:
4126:
4125:species group
4124:
4120:
4119:
4116:
4114:
4109:
4107:
4105:
4100:
4098:
4096:
4095:P. persephone
4090:
4089:species group
4088:
4084:
4083:
4080:
4078:
4073:
4071:
4069:
4064:
4062:
4060:
4055:
4053:
4051:
4050:P. brachyotis
4045:
4044:species group
4043:
4042:P. brachyotis
4039:
4038:
4036:
4026:
4025:
4020:
4014:
4012:
4007:
4005:
4003:
3998:
3996:
3994:
3989:
3987:
3985:
3980:
3979:
3977:
3975:
3974:
3969:
3963:
3961:
3956:
3954:
3952:
3947:
3946:
3944:
3934:
3933:
3928:
3922:
3920:
3915:
3913:
3911:
3906:
3904:
3902:
3897:
3895:
3893:
3888:
3886:
3884:
3879:
3877:
3875:
3870:
3868:
3866:
3861:
3860:
3858:
3856:
3855:
3850:
3844:
3842:
3837:
3835:
3833:
3828:
3827:
3825:
3823:
3822:
3817:
3811:
3809:
3804:
3802:
3800:
3795:
3794:
3792:
3782:
3781:
3776:
3770:
3768:
3763:
3761:
3759:
3754:
3753:
3751:
3749:
3748:
3743:
3737:
3735:
3730:
3728:
3726:
3721:
3719:
3717:
3712:
3710:
3708:
3703:
3702:
3700:
3698:
3697:
3692:
3686:
3684:
3679:
3677:
3675:
3670:
3668:
3666:
3661:
3659:
3657:
3652:
3650:
3648:
3643:
3641:
3639:
3634:
3632:
3630:
3625:
3623:
3621:
3616:
3614:
3612:
3607:
3605:
3603:
3598:
3596:
3594:
3589:
3587:
3585:
3580:
3578:
3576:
3571:
3570:
3568:
3558:
3557:
3552:
3546:
3544:
3539:
3538:
3536:
3534:
3533:
3528:
3525:
3519:
3511:
3507:
3503:
3499:
3493:
3489:
3471:
3469:
3464:
3463:
3461:
3459:
3458:
3453:
3447:
3445:
3440:
3439:
3437:
3435:
3434:
3429:
3426:
3424:
3420:
3410:
3408:
3403:
3401:
3399:
3394:
3392:
3390:
3385:
3383:
3381:
3376:
3374:
3372:
3371:P. albertisii
3367:
3366:
3364:
3362:
3361:
3360:Pseudochirops
3356:
3350:
3348:
3343:
3341:
3339:
3334:
3332:
3330:
3325:
3323:
3321:
3316:
3314:
3312:
3307:
3305:
3303:
3298:
3296:
3294:
3289:
3287:
3285:
3280:
3279:
3277:
3275:
3274:
3269:
3263:
3261:
3260:P. peregrinus
3256:
3255:
3253:
3251:
3250:
3249:Pseudocheirus
3245:
3239:
3237:
3232:
3231:
3229:
3227:
3226:
3221:
3215:
3213:
3208:
3206:
3204:
3199:
3197:
3195:
3194:P. armillatus
3190:
3189:
3187:
3177:
3176:
3171:
3165:
3163:
3162:H. lemuroides
3158:
3157:
3155:
3153:
3152:
3147:
3144:
3142:
3138:
3128:
3126:
3121:
3119:
3117:
3112:
3110:
3108:
3103:
3101:
3099:
3096:Biak glider (
3094:
3092:
3090:
3085:
3083:
3081:
3076:
3075:
3073:
3071:
3070:
3065:
3059:
3057:
3052:
3051:
3049:
3047:
3046:
3045:Gymnobelideus
3041:
3035:
3033:
3032:D. trivirgata
3028:
3026:
3024:
3019:
3017:
3015:
3010:
3008:
3006:
3001:
3000:
2998:
2996:
2995:
2990:
2987:
2985:
2981:
2971:
2969:
2964:
2963:
2961:
2959:
2958:
2953:
2950:
2948:
2944:
2934:
2932:
2927:
2925:
2923:
2918:
2916:
2914:
2909:
2907:
2905:
2900:
2899:
2897:
2895:
2894:
2889:
2883:
2881:
2876:
2875:
2873:
2871:
2870:
2865:
2862:
2852:
2848:
2838:
2836:
2831:
2830:
2828:
2826:
2825:
2820:
2814:
2812:
2807:
2805:
2803:
2802:T. johnstonii
2798:
2796:
2794:
2789:
2787:
2785:
2780:
2778:
2776:
2771:
2770:
2768:
2758:
2757:
2752:
2746:
2744:
2743:S. pelegensis
2739:
2737:
2735:
2734:S. celebensis
2730:
2729:
2727:
2725:
2724:
2719:
2713:
2711:
2706:
2704:
2702:
2697:
2695:
2693:
2688:
2686:
2684:
2679:
2677:
2675:
2670:
2669:
2667:
2665:
2664:
2659:
2653:
2651:
2646:
2644:
2642:
2637:
2635:
2633:
2628:
2626:
2624:
2619:
2617:
2615:
2614:P. orientalis
2610:
2608:
2606:
2601:
2599:
2597:
2592:
2590:
2588:
2583:
2581:
2579:
2574:
2572:
2570:
2565:
2563:
2561:
2556:
2554:
2552:
2551:P. carmelitae
2547:
2545:
2543:
2542:P. alexandrae
2540:Gebe cuscus (
2538:
2537:
2535:
2533:
2532:
2527:
2521:
2519:
2514:
2512:
2510:
2505:
2504:
2502:
2492:
2491:
2486:
2483:
2477:
2469:
2468:Phalangeridae
2465:
2461:
2457:
2448:
2444:
2426:
2424:
2419:
2417:
2415:
2410:
2409:
2407:
2405:
2404:
2399:
2393:
2391:
2386:
2385:
2383:
2381:
2380:
2375:
2372:
2362:
2358:
2348:
2346:
2341:
2340:
2338:
2336:
2335:
2334:Phascolarctos
2330:
2327:
2325:
2321:
2317:
2313:
2312:Vombatiformes
2307:
2303:
2297:
2293:
2291:
2287:
2285:
2281:
2279:
2275:
2274:
2271:
2266:
2265:Diprotodontia
2258:
2253:
2251:
2246:
2244:
2239:
2238:
2235:
2229:
2226:
2223:
2222:
2203:
2202:
2197:
2190:
2175:
2171:
2167:
2160:
2144:
2140:
2134:
2119:
2115:
2108:
2106:
2089:
2085:
2078:
2062:
2061:WWF-Australia
2058:
2052:
2036:
2032:
2025:
2010:
2006:
2000:
1985:. 15 May 2020
1984:
1980:
1974:
1958:
1954:
1948:
1932:
1928:
1922:
1907:. 2 July 2020
1906:
1902:
1896:
1890:
1885:
1877:
1873:
1868:
1863:
1859:
1855:
1851:
1850:in Australia"
1849:
1840:
1832:
1828:
1824:
1822:
1814:
1806:
1802:
1795:
1788:
1787:Claridge 2007
1783:
1767:
1763:
1759:
1755:
1749:
1747:
1740:, p. 66.
1739:
1738:Claridge 2007
1734:
1726:
1722:
1718:
1714:
1710:
1706:
1702:
1698:
1694:
1693:Vulpes vulpes
1690:
1686:
1679:
1672:
1671:Claridge 2007
1667:
1660:
1659:Claridge 2007
1655:
1636:
1632:
1628:
1621:
1619:
1611:
1604:
1603:Claridge 2007
1599:
1583:
1579:
1572:
1570:
1562:
1561:Claridge 2007
1557:
1541:
1537:
1530:
1528:
1526:
1524:
1515:
1511:
1507:
1503:
1499:
1495:
1488:
1481:
1480:Claridge 2007
1476:
1469:
1468:Claridge 2007
1464:
1456:
1452:
1447:
1442:
1438:
1434:
1430:
1427:(July 2001).
1426:
1422:
1416:
1409:
1408:Claridge 2007
1404:
1396:
1394:9780195573954
1390:
1386:
1382:
1378:
1372:
1370:
1368:
1366:
1357:
1353:
1346:
1342:
1336:
1334:
1332:
1330:
1328:
1326:
1309:
1303:
1295:
1289:
1281:
1274:
1266:
1260:
1245:
1238:
1231:
1229:
1220:
1216:
1212:
1206:
1202:
1201:
1196:
1195:Wilson, D. E.
1192:
1188:
1187:Groves, C. P.
1182:
1175:
1174:Claridge 2007
1170:
1154:
1153:
1148:
1142:
1134:
1130:
1128:
1119:
1117:
1101:
1097:
1091:
1075:
1070:
1066:
1062:
1061:
1056:
1054:
1045:
1043:
1041:
1036:
1029:
1027:9780643093416
1023:
1019:
1014:
1013:
1006:
1004:
1000:
995:
991:
987:
983:
979:
975:
971:
966:
964:
960:
955:
953:
949:
945:
940:
937:
933:
929:
925:
921:
916:
914:
910:
906:
902:
899:
895:
891:
887:
883:
878:
875:
871:
870:IUCN Red List
866:
863:
859:
855:
851:
847:
842:
840:
837:, run by the
836:
832:
822:
819:
813:
809:
803:
801:
796:
795:H. C. Richter
791:
787:
784:
780:
779:
774:
764:
762:
758:
754:
749:
744:
742:
738:
734:
733:Vulpes vulpes
730:
726:
722:
717:
715:
705:
703:
699:
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692:
690:
686:
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673:
672:
667:
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660:
659:fungal spores
656:
652:
651:
650:Hakea laurina
646:
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634:
633:
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625:As with the
618:
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502:Vulpes vulpes
499:
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246:
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198:
197:Binomial name
194:
190:
189:
184:
181:
180:
177:
176:
172:
169:
168:
165:
162:
159:
158:
155:
154:Diprotodontia
152:
149:
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139:
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135:
132:
129:
128:
125:
122:
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118:
115:
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109:
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105:
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99:
98:
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84:
78:
72:
67:
61:
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41:
36:
32:
27:
22:
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4882:
4828:
4567:
4529:H. moschatus
4528:
4516:
4492:P. gilbertii
4491:
4482:
4473:
4453:
4441:
4432:
4430:
4423:
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4394:
4383:A. rufescens
4382:
4370:
4345:
4333:
4321:
4312:
4303:
4294:
4285:
4276:
4267:
4256:(pademelons)
4247:
4235:
4223:
4211:
4202:
4193:
4184:
4176:P. lateralis
4175:
4166:
4157:
4148:
4140:P. coenensis
4139:
4131:P. assimilis
4130:
4122:
4121:
4113:P. xanthopus
4112:
4103:
4094:
4087:P. xanthopus
4086:
4085:
4076:
4067:
4059:P. burbidgei
4058:
4049:
4041:
4040:
4022:
4010:
4001:
3993:O. bernardus
3992:
3983:
3971:
3960:O. unguifera
3959:
3950:
3930:
3918:
3909:
3900:
3891:
3882:
3873:
3864:
3854:Notamacropus
3852:
3841:M. giganteus
3840:
3831:
3819:
3807:
3798:
3780:Lagorchestes
3778:
3767:D. vanheurni
3766:
3757:
3745:
3733:
3724:
3715:
3706:
3694:
3682:
3674:D. stellarum
3673:
3664:
3655:
3646:
3637:
3628:
3620:D. matschiei
3619:
3611:D. lumholtzi
3610:
3601:
3592:
3583:
3574:
3554:
3543:L. fasciatus
3542:
3532:Lagostrophus
3530:
3510:Macropodidae
3467:
3457:Distoechurus
3455:
3443:
3431:
3406:
3398:P. coronatus
3397:
3388:
3379:
3370:
3358:
3347:P. schlegeli
3346:
3337:
3328:
3319:
3310:
3301:
3292:
3284:P. canescens
3283:
3271:
3259:
3247:
3235:
3225:Petropseudes
3223:
3211:
3202:
3193:
3173:
3161:
3151:Hemibelideus
3149:
3124:
3115:
3107:P. breviceps
3106:
3098:P. biacensis
3097:
3089:P. australis
3088:
3079:
3067:
3055:
3043:
3031:
3022:
3013:
3004:
2994:Dactylopsila
2992:
2968:T. rostratus
2967:
2955:
2947:Tarsipedidae
2930:
2921:
2913:C. concinnus
2912:
2903:
2891:
2879:
2867:
2834:
2822:
2811:T. vulpecula
2810:
2801:
2792:
2783:
2774:
2754:
2742:
2733:
2723:Strigocuscus
2721:
2709:
2701:S. rufoniger
2700:
2692:S. papuensis
2691:
2683:S. maculatus
2682:
2673:
2661:
2649:
2640:
2631:
2622:
2613:
2604:
2595:
2586:
2577:
2568:
2559:
2550:
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2529:
2517:
2509:A. melanotis
2508:
2488:
2422:
2414:L. latifrons
2413:
2401:
2389:
2377:
2344:
2332:
2205:. Retrieved
2199:
2189:
2177:. Retrieved
2169:
2159:
2147:. Retrieved
2145:. 2 May 2022
2142:
2133:
2121:. Retrieved
2117:
2092:. Retrieved
2087:
2077:
2065:. Retrieved
2060:
2051:
2039:. Retrieved
2034:
2024:
2012:. Retrieved
2008:
1999:
1987:. Retrieved
1982:
1973:
1961:. Retrieved
1956:
1947:
1935:. Retrieved
1930:
1921:
1909:. Retrieved
1904:
1895:
1884:
1857:
1853:
1847:
1839:
1820:
1813:
1804:
1794:
1782:
1772:16 September
1770:. Retrieved
1766:the original
1757:
1733:
1703:(1): 18–27.
1700:
1696:
1695:) control".
1692:
1688:
1684:
1678:
1666:
1654:
1642:. Retrieved
1635:the original
1626:
1617:
1610:
1598:
1586:. Retrieved
1581:
1556:
1546:23 September
1544:. Retrieved
1539:
1497:
1493:
1487:
1475:
1463:
1436:
1432:
1415:
1410:, p. 3.
1403:
1384:
1355:
1351:
1314:12 September
1312:. Retrieved
1302:
1279:
1273:
1247:. Retrieved
1243:
1199:
1181:
1176:, p. 5.
1169:
1157:. Retrieved
1150:
1141:
1132:
1126:
1103:. Retrieved
1099:
1090:
1080:25 September
1078:. Retrieved
1064:
1058:
1052:
1017:
994:Wedge Island
990:translocated
967:
956:
941:
917:
879:
867:
854:Wedge Island
843:
828:
825:Conservation
814:
810:
806:
798:
776:
770:
745:
740:
732:
725:Aquila audax
724:
718:
711:
695:
693:
675:
669:
663:
648:
630:
624:
608:
599:fire regimes
596:
593:
588:
558:
551:
544:John Gilbert
537:
530:
526:Duisburg Zoo
506:
501:
451:
435:
427:
425:
405:
404:
390:
377:
373:
369:
361:
359:
341:
337:
333:
327:
322:
307:
297:
293:
268:
267:
263:
259:
257:
234:
228:
205:
203:
187:
186:
174:
140:Infraclass:
18:
4695:iNaturalist
4592:Wikispecies
4474:P. longipes
4415:B. gaimardi
4372:Aepyprymnus
4212:P. sharmani
4167:P. inornata
4158:P. herberti
4077:P. wilkinsi
4068:P. concinna
4002:O. robustus
3951:O. fraenata
3932:Onychogalea
3874:N. dorsalis
3808:L. hirsutus
3758:D. macleayi
3747:Dorcopsulus
3734:D. muelleri
3725:D. luctuosa
3584:D. dorianus
3556:Dendrolagus
3468:D. pennatus
3444:A. pygmaeus
3423:Acrobatidae
3389:P. corinnae
3329:P. larvatus
3302:P. cinereus
3175:Petauroides
3116:P. gracilis
3014:D. palpator
3005:D. megalura
2904:C. caudatus
2893:Cercartetus
2851:Burramyidae
2756:Trichosurus
2674:S. kraemeri
2663:Spilocuscus
2650:P. vestitus
2641:P. sericeus
2587:P. matabiru
2578:P. lullulae
2560:P. gymnotis
2474:(including
2423:L. krefftii
2403:Lasiorhinus
2345:P. cinereus
2296:Marsupialia
2294:Infraclass
2094:3 September
2014:3 September
1989:3 September
1963:22 February
1911:3 September
1805:7.30 report
1685:Felis catus
1358:(3): 42–48.
909:understorey
898:sclerophyll
797:in Gould's
741:Felis catus
676:Mesophellia
671:Mesophellia
637:fungivorous
573:agriculture
569:pastoralism
401:Description
221:Subspecies
144:Marsupialia
4955:Categories
4462:(potoroos)
4442:B. tropica
4424:B. lesueur
4403:(bettongs)
4362:Potoroidae
4346:W. bicolor
4304:T. lanatus
4295:T. calabyi
4185:P. mareeba
4149:P. godmani
4066:Nabarlek (
3973:Osphranter
3883:N. eugenii
3683:D. ursinus
3665:D. scottae
3638:D. notatus
3602:D. inustus
3516:(includes
3407:P. cupreus
3380:P. archeri
3311:P. forbesi
2984:Petauridae
2922:C. lepidus
2784:T. caninus
2710:S. wilsoni
2623:P. ornatus
2605:P. mimicus
2596:P. matanim
2518:A. ursinus
2390:V. ursinus
2361:Vombatidae
1425:Calaby, J.
1381:Knight, F.
1341:Abbott, I.
1105:14 January
1009:References
894:feral cats
689:methionine
629:and other
524:Woylie at
464:, most of
458:south-west
446:See also:
350:John Gould
310:J. E. Gray
278:Potoroidae
213:J. E. Gray
164:Potoroidae
73:Appendix I
4966:Bettongia
4898:Q27828377
4844:Q27828380
4396:Bettongia
4322:T. thetis
4286:T. brunii
4277:T. browni
4249:Thylogale
4024:Petrogale
3910:N. parryi
3865:N. agilis
3716:D. hageni
3707:D. atrata
3696:Dorcopsis
3663:Tenkile (
3656:D. spadix
3629:D. mbaiso
3627:Dingiso (
3518:wallabies
3496:Suborder
3433:Acrobates
3338:P. mayeri
3293:P. caroli
3212:P. volans
2880:B. parvus
2531:Phalanger
2455:(possums)
2451:Suborder
2367:(wombats)
2310:Suborder
2009:Shark Bay
1876:0030-6053
1823:- Woylie"
1717:1446-5698
1455:0067-2238
1421:Short, J.
1100:cites.org
932:Tutanning
884:south of
858:Shark Bay
846:Venus Bay
831:red foxes
773:nocturnal
767:Behaviour
748:ecosystem
737:feral cat
643:, seeds,
632:Bettongia
581:feral cat
509:potoroine
410:potoroine
286:marsupial
182:Species:
175:Bettongia
110:Kingdom:
104:Eukaryota
4892:Wikidata
4838:Wikidata
4812:Species+
4765:11000182
4713:10453525
4577:Wikidata
4455:Potorous
4431:Woylie (
4422:Boodie (
4335:Wallabia
4234:Quokka (
4057:Monjon (
4011:O. rufus
3901:N. parma
3821:Macropus
3236:P. dahli
3203:P. minor
3080:P. abidi
3069:Petaurus
3023:D. tatei
2957:Tarsipes
2931:C. nanus
2869:Burramys
2490:Ailurops
2476:cuscuses
2379:Vombatus
2290:Mammalia
2284:Chordata
2278:Animalia
2276:Kingdom
2179:3 August
2170:ABC News
2123:3 August
2088:ABC News
2035:ABC News
1937:6 August
1889:Wadderin
1725:84854873
1582:ABC News
1540:ABC News
1383:(2011).
1343:(2008).
1259:cite web
1249:25 April
1219:62265494
1189:(2005).
1159:13 March
948:Newhaven
934:and the
874:parasite
818:hectares
761:bushland
685:cysteine
655:bacteria
627:Potoroos
597:Changed
492:and the
470:Victoria
454:spinifex
314:holotype
304:Taxonomy
160:Family:
134:Mammalia
124:Chordata
120:Phylum:
114:Animalia
100:Domain:
60:IUCN 3.1
4752:1000323
4687:2440026
4583:Q210070
4225:Setonix
3892:N. irma
3636:Ifola (
2343:Koala (
2282:Phylum
2267:species
2263:Extant
2207:26 June
2067:6 April
2041:6 April
1588:6 April
1502:Bibcode
999:Noongar
965:in WA.
729:red fox
708:Ecology
645:insects
605:Current
565:grazing
498:red fox
494:Denmark
364:in the
170:Genus:
150:Order:
130:Class:
75: (
58: (
24:Woylie
4925:881300
4804:231481
4726:552701
4674:128419
4658:ECOS:
4638:718688
4620:ARKive
2824:Wyulda
2288:Class
1874:
1723:
1715:
1644:19 May
1453:
1391:
1217:
1207:
1024:
901:forest
890:numbat
802:, 1863
753:spores
735:) and
681:lycine
641:tubers
415:pelage
387:boodie
378:woylie
274:mammal
260:woylie
215:, 1837
4938:66844
4933:SPRAT
4871:66658
4866:SPRAT
4791:97768
4778:69259
4708:IRMNG
4700:42998
4651:68JYM
2149:2 May
2143:7NEWS
1721:S2CID
1638:(PDF)
1623:(PDF)
1348:(PDF)
1240:(PDF)
1193:. In
992:from
928:Perup
886:Perth
783:nests
757:fungi
702:scats
480:, by
362:walyu
282:fungi
77:CITES
71:CITES
4920:NCBI
4817:6709
4773:NCBI
4739:2785
4734:IUCN
4721:ITIS
4682:GBIF
4661:5516
4633:BOLD
2209:2023
2181:2022
2151:2022
2125:2022
2096:2020
2069:2021
2043:2021
2016:2020
1991:2020
1965:2022
1939:2021
1913:2020
1872:ISSN
1854:Oryx
1774:2011
1713:ISSN
1646:2014
1590:2021
1548:2019
1451:ISSN
1389:ISBN
1316:2012
1294:link
1265:link
1251:2022
1215:OCLC
1205:ISBN
1161:2019
1107:2022
1082:2021
1065:2016
1022:ISBN
903:and
852:and
687:and
621:Diet
579:and
571:and
376:and
374:woli
258:The
4907:AFD
4853:AFD
4760:MSW
4747:MDD
4669:EoL
4646:CoL
4607:AFD
1862:doi
1705:doi
1510:doi
1441:doi
1069:doi
911:of
755:of
460:of
370:wol
262:or
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