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Amphidromus perversus

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506: 454: 476: 757: 33: 911: 42: 63: 1121: 842:; Djukung specimens are in the Zoological Museum in Amsterdam; and topotypes from Bajutan are in Chicago Natural History Museum, no. 97806, and in Butot and the Zoologisch Museum, Amsterdam, Some additional material from Kangean Island (USNM 468416, Paravicini!) was seen after the description had been written. 441:
The shell is solid, polished, amphidromine, moderately large (45–55 mm) high, usually with a dark varix. Lip and callus white; ground color white, orange, yellow, or yellowish-green. Many specimens with interrupted or continuous radial brownish streaks covering part or all of the whorls on the
790:
Remarks: The problem of how to treat insular populations which show minor divergence from the main stocks is far from being satisfactorily settled. The intense coloration of the Kangean populations shows a minor difference from the main populations and Laidlaw & Solem (1961) have recognized it
1171:
Linnaeus, C. (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata [10th revised edition], vol. 1: 824 pp. Laurentius Salvius:
786:
Description: A brilliantly colored series of populations with highly polished shell surface. The general intensity of the coloration and the strong polish are the only features separating this race from the other subspecies.
744:
Lamarck, 1822, has wavy brown bands running over the entire whorl. The bands are interrupted in the middle of the whorl by a narrow spiral band of the ground color. This variation is found in all parts of the species
976:
This weakly characterized race differs primarily by its intense color, strong color zone, and slight white margin at the suture. Of seventeen shells collected in May, 1954, by Hoogerwerf at
835: 772: 855:
Remarks: The shell is white with rather numerous brown flammulations that become confluent on the base, elongate, and quite slender. Apparently it is a white derivative from the
791:
with a subspecific name. For purposes of future reference, they selected as holotype a shell with a color pattern quite distinct from any which have been named. Thus, the name
980:, Bawean Island, ten were dextral and seven sinistral. Three lacked a varix, one was grass green in color, three were whitish, and thirteen had yellow ground color. The name 892:, the numerous wavy brown flammulations that are partially interrupted by a yellow peripheral color band, and the solid shell are characteristic. This variety is similar to 900:
voyage is questionable, and it is quite possible that the Perak records of De Morgan are in error. The only certain localities are the Singapore area and Riau Archipelago.
642:. Instead of flammulations and a basal purplish zone it has a pale green or greenish-yellow base which is only slightly darker than the greenish-yellow or citron-colored 939:
is quite variable in color, ranging from white to dark (see Pilsbry 1900, pp. 162–163). Possibly some of the material that Jacobi (1895) dissected under the names
534:
Dillwyn, 1817 (not Martyn, 1784), is usable for the orange-colored (instead of yellow) monochrome shells found on Java and Celebes. Apparently an individual variation.
734:
von Martens, 1867, has continuous, rather narrow, straight brown bands running from the suture to the bottom of the whorl. It is known from Bali, Borneo, and Celebes.
599: 921: 614:
from East Borneo. The flammulations above the periphery are reduced in number. Possibly this may prove to be a subspecies, but not enough data are available.
558: 577: 947:
belongs here, but the shells were not figured and their location is unknown. The South Natuna Island record needs confirmation, while specimens from
724:
Linnaeus, 1758, is a solid yellow shell with black varix and narrow black band behind the peristome. It is known from the whole range of the species.
385: 977: 882: 594:
Pilsbry, 1900, is based on a stout variant of the interruptus pattern, whitish in background color, and rather similar to the Javanese subspecies,
549: 381: 568:
complex and the lower parts of the whorls are yellow, with or without narrow, wavy, reddish-brown flammulations. The original specimens in the
1286: 1182: 827: 562:
Butot, is quite different and the actual locahty of aureus is still unknown. The shells have a wide, white zone below the suture as in the
1007:
MolluscaBase eds. (2020). MolluscaBase. Amphidromus perversus (Linnaeus, 1758). Accessed through: World Register of Marine Species at:
948: 860: 776: 1260: 1154:
Schileyko, A. A. (2011). Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica. 21 (1): 1-68
896:, but has only partial interruption of the stripes by the color band and has the purple apex. The Biliton record from the early 1358: 1363: 795:
can at least be used for the color variation if it is decided to include the Kangean populations in the nominate form of
544:
Martyn, 1784 — It still have not been reported from a precise locality. It was based on three shells collected by Sir
628:
Müller, 1774. At best it might be considered a slightly different stage in an almost continuous series of variations.
1291: 696:
P. & F. Sarasin, 1899 is snow-white with a black varix and black mark behind the peristome. It is an "albino"
292: 178: 831: 710:
von Martens, 1867, is an unfigured variety which probably can be equated with squat individuals of the typical
603: 569: 1330: 62: 1160:
Rensch, B. (1938). Neue Landschnecken von der Insel Penida. Zoologischer Anzeiger. 123: 10/12: 302-306.
1317: 1176: 1008: 1157:
Vermeulen, J.J. & Whitten, A.J. (1998). Guide to the land snails of Bali, 164. Leiden: Backhuys.
984:
Fruhstorfer, 1905, refers to white shells with a very dark color band. This variation is similar to
863:, East Java (Butot, Chicago Natural History Museum 72404) ranged from 43.5 to 48 mm in height. 505: 564: 498:
undoubtedly offers the greatest array of discretely variable color combinations of any species of
453: 1124: 664:, and brownish flammulations that are somewhat interrupted on the periphery and do not reach the 434:
species, the lip is internally thickened, forms a "roll" in its expansion, and has a very heavy
1202: 660:
Müller, 1774, has a purplish or brownish zone on the base, a yellow or whitish patch near the
475: 1304: 661: 157: 1353: 1247: 910: 576:, but the locality remains a mystery. Note: Martyn's work was invalidated by action of the 826:
Paratypes: Specimens were seen from several localities on Kangean Island. The shells from
678:
von Martens, 1867, is an unfigured variety from Bali and seems to be intermediate between
8: 1020: 859:
stock, which may have a definite geographic separation in Bali and Java. Material from
665: 264: 233: 187: 57: 1132: 1299: 1234: 756: 874: 1239: 32: 780: 643: 435: 405: 114: 423:
are amphidromine (left-handed and right-handed snails occur in the population).
1225: 955:(CNHM 72427), are referred by Laidlaw & Solem (1961) with some hesitation. 952: 889: 1170: 41: 1347: 966: 288: 174: 545: 409: 397: 256: 700:
which is common in the Celebes and sporadically seen from Java and Borneo.
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von Martens, 1867, was considered by Pilsbry (1900) to be identical with
134: 1265: 1128: 443: 246: 229: 124: 104: 1278: 1188: 607: 393: 239: 236: 74: 1196: 1219: 897: 761: 458: 370: 94: 1009:
http://www.marinespecies.org/aphia.php?p=taxdetails&id=1026814
1325: 996:
This article incorporates public domain text from the reference
389: 358: 242: 225: 1252: 611: 401: 366: 84: 638:
von Martens, 1867, probably is a derivative of the Celebesian
580:, varietal names are not accorded legal status and the use of 1312: 878: 839: 970: 932: 374: 362: 1120: 392:
needs confirmation. Probably it has been introduced into
924:; possibly the South Natuna and Anamba Islands also. 958: 903: 578:
International Commission on Zoological Nomenclature
866: 845: 1345: 1115: 1113: 1111: 1109: 1107: 1105: 1103: 1101: 1099: 1097: 1095: 1093: 1091: 1089: 1087: 1085: 1083: 1081: 1079: 1077: 1075: 1073: 1071: 1069: 1067: 1065: 1063: 1061: 1059: 1057: 1055: 1053: 749: 49:Abapertural and apertural view of the shell of 1051: 1049: 1047: 1045: 1043: 1041: 1039: 1037: 1035: 1033: 783:. Collected by Hoogerwerf on August 20, 1954. 1183:National Center for Biotechnology Information 852:Range: Central and East Java, possibly Bali. 598:. It was described from South Borneo in the 1030: 479:apertural view of the shell of paratype of 931:is absent; otherwise this is close to the 40: 31: 909: 799:. Besides the nominate color pattern of 755: 650:but differs in the darker basal portion. 504: 474: 452: 606:63377), but there are specimens in the 1346: 396:. Subspecies recognized here are from 1201: 1200: 823:pattern were seen on Kangean Island. 584:is as a convenient term of reference. 1331:9DDCCD77-E9CE-49F3-891C-FE3909D29B69 1318:768024CA-51CE-E4F3-416F-44D0C51983B2 1001: 415: 263:, by the subsequent designation of 197:Amphidromus (Amphidromus) perversus 13: 202:accepted, alternate representation 14: 1375: 1164: 960:Amphidromus perversus rufocinctus 332:Amphidromus perversus rufocinctus 1178:Amphidromus perversus natunensis 1119: 937:Amphidromus perversus natunensis 915:Amphidromus perversus natunensis 905:Amphidromus perversus natunensis 646:. It is most similar to typical 324:Amphidromus perversus natunensis 61: 868:Amphidromus perversus melanomma 847:Amphidromus perversus emaciatus 760:apertural view of the shell of 738:Amphidromus perversus perversus 728:Amphidromus perversus perversus 718:Amphidromus perversus perversus 704:Amphidromus perversus perversus 690:Amphidromus perversus perversus 672:Amphidromus perversus perversus 654:Amphidromus perversus perversus 632:Amphidromus perversus perversus 618:Amphidromus perversus perversus 588:Amphidromus perversus perversus 538:Amphidromus perversus perversus 528:Amphidromus perversus perversus 521:Amphidromus perversus perversus 510:Amphidromus perversus perversus 457:apertural view of the shell of 348: 316:Amphidromus perversus melanomma 308:Amphidromus perversus emaciatus 284:Amphidromus perversus perversus 1014: 927:The yellow peripheral band of 836:Zoological Museum in Amsterdam 832:Chicago Natural History Museum 773:Zoological Museum in Amsterdam 604:Chicago Natural History Museum 340:Amphidromus perversus siglerae 1: 991: 430:and most other thick-shelled 270: 1359:Gastropods described in 1758 986:Amphidromus perversus butoti 877:near Singapore and possibly 766:Amphidromus perversus butoti 751:Amphidromus perversus butoti 481:Amphidromus perversus butoti 463:Amphidromus perversus butoti 300:Amphidromus perversus butoti 275:There are six subspecies of 7: 1364:Taxa named by Carl Linnaeus 10: 1380: 803:, shells referable to the 1209: 303:Laidlaw & Solem, 1961 193: 186: 163: 156: 58:Scientific classification 56: 48: 39: 30: 23: 565:Amphidromus atricallosus 1137:: a synoptic catalogue" 941:Amphidromus interruptus 775:. The type locality is 572:appear to be a form of 1133:"The land snail genus 917: 768: 600:Bandjermassin District 570:Natural History Museum 516: 488: 470: 211:(original combination) 1211:Amphidromus perversus 1191:Amphidromus perversus 913: 759: 574:Amphidromus perversus 508: 496:Amphidromus perversus 478: 456: 428:Amphidromus perversus 421:Amphidromus perversus 355:Amphidromus perversus 277:Amphidromus perversus 253:Amphidromus perversus 221:Amphidromus perversus 167:Amphidromus perversus 51:Amphidromus perversus 25:Amphidromus perversus 1189:photos of shells of 922:Great Natuna Islands 888:Remarks: The purple 834:, no. 97808, in the 388:. Its occurrence on 16:Species of gastropod 945:Amphidromus chloris 519:Color variation of 311:(von Martens, 1867) 918: 769: 559:Amphidromus banksi 523:include 12 forms: 517: 489: 471: 265:Eduard von Martens 1341: 1340: 1300:Open Tree of Life 1203:Taxon identifiers 1141:Fieldiana Zoology 493: 492: 416:Shell description 344: 336: 335:Fruhstorfer, 1905 328: 320: 312: 304: 296: 228:of air-breathing 217: 216: 210: 201: 200:(Linnaeus, 1758)· 149:A. perversus 1371: 1334: 1333: 1321: 1320: 1308: 1307: 1295: 1294: 1282: 1281: 1269: 1268: 1256: 1255: 1243: 1242: 1230: 1229: 1228: 1198: 1197: 1147: 1123: 1117: 1028: 1027:2nd ed., p. 184. 1018: 1012: 1005: 875:Riau Archipelago 548:, supposedly on 449: 448: 442:lower spire and 380:Absent from the 353:Distribution of 342: 334: 326: 319:(Pfeiffer, 1852) 318: 310: 302: 286: 208: 199: 169: 66: 65: 44: 35: 21: 20: 1379: 1378: 1374: 1373: 1372: 1370: 1369: 1368: 1344: 1343: 1342: 1337: 1329: 1324: 1316: 1311: 1303: 1298: 1290: 1285: 1277: 1272: 1264: 1259: 1251: 1246: 1238: 1233: 1224: 1223: 1218: 1205: 1167: 1151: 1150: 1118: 1031: 1019: 1015: 1006: 1002: 994: 963: 908: 871: 850: 781:Kangean Islands 754: 436:parietal callus 418: 406:Kangean Islands 386:Panaitan Island 351: 273: 182: 171: 165: 152: 115:Stylommatophora 60: 17: 12: 11: 5: 1377: 1367: 1366: 1361: 1356: 1339: 1338: 1336: 1335: 1322: 1309: 1296: 1283: 1270: 1257: 1244: 1231: 1215: 1213: 1207: 1206: 1195: 1194: 1186: 1174: 1166: 1165:External links 1163: 1162: 1161: 1158: 1155: 1149: 1148: 1029: 1021:von Martens E. 1013: 999: 998: 993: 990: 978:Telaga Kastoba 962: 957: 953:Anamba Islands 907: 902: 883:Biliton Island 870: 865: 849: 844: 753: 748: 747: 746: 735: 725: 715: 701: 687: 669: 651: 629: 615: 585: 550:Pulau Panaitan 535: 491: 490: 487:color pattern. 472: 469:color pattern. 417: 414: 350: 347: 346: 345: 337: 329: 321: 313: 305: 297: 272: 269: 245:in the family 215: 214: 213: 212: 209:Linnaeus, 1758 206:Helix perversa 203: 191: 190: 184: 183: 172: 161: 160: 154: 153: 146: 144: 140: 139: 132: 128: 127: 122: 118: 117: 112: 108: 107: 102: 98: 97: 92: 88: 87: 82: 78: 77: 72: 68: 67: 54: 53: 46: 45: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 1376: 1365: 1362: 1360: 1357: 1355: 1352: 1351: 1349: 1332: 1327: 1323: 1319: 1314: 1310: 1306: 1301: 1297: 1293: 1288: 1284: 1280: 1275: 1271: 1267: 1262: 1258: 1254: 1249: 1245: 1241: 1236: 1232: 1227: 1221: 1217: 1216: 1214: 1212: 1208: 1204: 1199: 1193: 1192: 1187: 1184: 1180: 1179: 1175: 1173: 1169: 1168: 1159: 1156: 1153: 1152: 1146:(4): 505-720. 1145: 1142: 1138: 1136: 1130: 1126: 1125:Laidlaw F. F. 1122: 1116: 1114: 1112: 1110: 1108: 1106: 1104: 1102: 1100: 1098: 1096: 1094: 1092: 1090: 1088: 1086: 1084: 1082: 1080: 1078: 1076: 1074: 1072: 1070: 1068: 1066: 1064: 1062: 1060: 1058: 1056: 1054: 1052: 1050: 1048: 1046: 1044: 1042: 1040: 1038: 1036: 1034: 1026: 1022: 1017: 1011:on 2020-05-17 1010: 1004: 1000: 997: 989: 987: 983: 979: 974: 972: 968: 967:Bawean Island 961: 956: 954: 950: 946: 942: 938: 934: 930: 925: 923: 916: 912: 906: 901: 899: 895: 891: 886: 884: 880: 876: 869: 864: 862: 858: 853: 848: 843: 841: 837: 833: 829: 824: 822: 818: 814: 810: 806: 802: 798: 794: 788: 784: 782: 778: 774: 767: 763: 758: 752: 743: 739: 736: 733: 729: 726: 723: 719: 716: 713: 709: 705: 702: 699: 695: 691: 688: 685: 681: 677: 673: 670: 667: 663: 659: 655: 652: 649: 645: 641: 637: 633: 630: 627: 623: 619: 616: 613: 609: 605: 601: 597: 593: 589: 586: 583: 579: 575: 571: 567: 566: 561: 560: 556:found there, 555: 551: 547: 543: 539: 536: 533: 529: 526: 525: 524: 522: 515: 511: 507: 503: 501: 497: 486: 482: 477: 473: 468: 464: 460: 455: 451: 450: 447: 445: 439: 437: 433: 429: 424: 422: 413: 411: 407: 403: 399: 395: 391: 387: 383: 382:Mentawi Chain 378: 376: 372: 368: 364: 360: 356: 341: 338: 333: 330: 325: 322: 317: 314: 309: 306: 301: 298: 294: 290: 285: 282: 281: 280: 278: 268: 266: 262: 259:of the genus 258: 254: 250: 248: 244: 241: 238: 235: 231: 227: 223: 222: 207: 204: 198: 195: 194: 192: 189: 185: 180: 176: 170: 168: 162: 159: 158:Binomial name 155: 151: 150: 145: 142: 141: 138: 137: 133: 130: 129: 126: 123: 120: 119: 116: 113: 110: 109: 106: 103: 100: 99: 96: 93: 90: 89: 86: 83: 80: 79: 76: 73: 70: 69: 64: 59: 55: 52: 47: 43: 38: 34: 29: 26: 22: 19: 1210: 1190: 1177: 1143: 1140: 1134: 1025:Die Heliceen 1024: 1016: 1003: 995: 985: 981: 975: 964: 959: 944: 940: 936: 935:population. 928: 926: 919: 914: 904: 893: 887: 885:off Borneo. 872: 867: 856: 854: 851: 846: 825: 820: 819:and typical 816: 812: 808: 805:infraviridis 804: 800: 796: 792: 789: 785: 770: 765: 750: 741: 737: 731: 727: 721: 717: 711: 707: 703: 697: 693: 689: 683: 679: 675: 671: 657: 653: 647: 639: 636:infraviridis 635: 631: 625: 621: 617: 595: 591: 587: 581: 573: 563: 557: 553: 546:Joseph Banks 541: 537: 531: 527: 520: 518: 513: 509: 499: 495: 494: 484: 480: 467:infraviridis 466: 462: 440: 431: 427: 425: 420: 419: 410:Riau Islands 398:Great Natuna 379: 354: 352: 349:Distribution 339: 331: 327:Fulton, 1896 323: 315: 307: 299: 283: 276: 274: 260: 257:type species 252: 251: 220: 219: 218: 205: 196: 166: 164: 148: 147: 135: 50: 24: 18: 1354:Amphidromus 1274:iNaturalist 1135:Amphidromus 857:interruptus 813:rufocinctus 809:infrapictus 680:interruptus 658:interruptus 640:infrapictus 626:interruptus 622:infrapictus 554:Amphidromus 552:. The only 500:Amphidromus 432:Amphidromus 343:Thach, 2018 261:Amphidromus 234:terrestrial 136:Amphidromus 1348:Categories 992:References 982:sankapurus 828:Sepandjang 771:Holotype: 592:borneensis 444:body whorl 271:Subspecies 247:Camaenidae 230:land snail 125:Camaenidae 105:Gastropoda 929:melanomma 838:, and in 821:perversus 797:perversus 732:strigosus 722:perversus 712:perversus 698:perversus 662:columella 648:perversus 608:Naturalis 596:emaciatus 394:Singapore 240:gastropod 237:pulmonate 143:Species: 81:Kingdom: 75:Eukaryota 1226:Q4748268 1220:Wikidata 1131:(1961). 1129:Solem A. 1023:(1860). 898:Samarang 894:sultanus 817:sultanus 762:holotype 742:sultanus 714:pattern. 684:sultanus 485:sultanus 483:showing 465:showing 459:paratype 371:Sulawesi 357:include 289:Linnaeus 267:(1860). 188:Synonyms 175:Linnaeus 121:Family: 95:Mollusca 91:Phylum: 85:Animalia 71:Domain: 1326:ZooBank 1305:7500440 1292:2202199 1266:8823723 1253:4895551 1172:Holmiae 965:Range: 949:Siantan 920:Range: 873:Range: 861:Kedewan 830:are in 777:Bajutan 390:Sumbawa 359:Sumatra 255:is the 243:mollusc 226:species 131:Genus: 111:Order: 101:Class: 1279:509466 1185:(NCBI) 1127:& 801:butoti 793:butoti 745:range. 708:obesus 694:niveus 668:above. 666:suture 612:Leiden 582:aureus 542:aureus 514:niveus 402:Bawean 367:Borneo 1313:Plazi 879:Perak 840:Butot 676:mitra 644:spire 532:aurea 224:is a 1287:NCBI 1261:GBIF 1240:D2N5 971:Java 943:and 933:Riau 890:apex 881:and 682:and 408:and 384:and 375:Bali 373:and 363:Java 361:and 293:1758 232:, a 179:1758 1248:EoL 1235:CoL 1181:at 764:of 740:f. 730:f. 720:f. 706:f. 692:f. 674:f. 656:f. 634:f. 620:f. 590:f. 540:f. 530:f. 512:f. 461:of 426:In 365:to 1350:: 1328:: 1315:: 1302:: 1289:: 1276:: 1263:: 1250:: 1237:: 1222:: 1144:41 1139:. 1032:^ 988:. 973:. 969:, 951:, 815:, 811:, 807:, 779:, 610:, 502:. 446:. 438:. 412:. 404:, 400:, 377:. 369:, 291:, 279:: 249:. 177:, 686:. 602:( 295:) 287:( 181:) 173:(

Index



Scientific classification
Edit this classification
Eukaryota
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Amphidromus
Binomial name
Linnaeus
1758
Synonyms
species
land snail
terrestrial
pulmonate
gastropod
mollusc
Camaenidae
type species
Eduard von Martens
Linnaeus
1758
Sumatra
Java
Borneo
Sulawesi

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