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Basidiomycota

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compatible nuclei following meiosis migrate into each basidiospore, which is then dispersed as a pre-existing dikaryon. Often such species form only two spores per basidium, but that too varies. Following meiosis, mitotic divisions can occur in the basidium. Multiple numbers of basidiospores can result, including odd numbers via degeneration of nuclei, or pairing up of nuclei, or lack of migration of nuclei. For example, the chanterelle genus
367:) outside of these, among the Basidiomycota. As now classified, the subphyla join and also cut across various obsolete taxonomic groups (see below) previously commonly used to describe Basidiomycota. According to a 2008 estimate, Basidiomycota comprise three subphyla (including six unassigned classes) 16 classes, 52 orders, 177 families, 1,589 genera, and 31,515 species. Wijayawardene et al. 2020 produced an update that recognized 19 classes ( 1157:. can have one-, two-, three- or four-spored basidia under some circumstances. Occasionally, monokaryons of some taxa can form morphologically fully formed basidiomes and anatomically correct basidia and ballistic basidiospores in the absence of dikaryon formation, diploid nuclei, and meiosis. A rare few number of taxa have extended diploid lifecycles, but can be common species. Examples exist in the mushroom genera 1065:
multiple compatibility factors. Plasmogamy between compatible individuals leads to delayed karyogamy leading to establishment of a dikaryon. The dikaryon is long lasting but ultimately gives rise to either fruitbodies with basidia or directly to basidia without fruitbodies. The paired dikaryon in the basidium fuse (i.e. karyogamy takes place). The diploid basidium begins the cycle again.
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which are tapered spine-like projections on basidia, and are typically curved, like the horns of a bull. In some Basidiomycota the spores are not ballistic, and the sterigmata may be straight, reduced to stubs, or absent. The basidiospores of these non-ballistosporic basidia may either bud off, or be
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Padamsee, M.; Kumar, T. K.; Riley, R.; Binder, M.; Boyd, A.; Calvo, A. M.; Furukawa, K.; Hesse, C.; Hohmann, S.; James, T. Y.; LaButti, K.; Lapidus, A.; Lindquist, E.; Lucas, S.; Miller, K.; Shantappa, S.; Grigoriev, I. V.; Hibbett, D. S.; McLaughlin, D. J.; Spatafora, J. W.; Aime, M. C. (Mar 2012).
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and resultant monokaryons, have nuclei that are compatible with 50% (if bipolar) or 25% (if tetrapolar) of their sister basidiospores (and their resultant monokaryons) because the mating genes must differ for them to be compatible. However, there are sometimes more than two possible alleles for a
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In summary, meiosis takes place in a diploid basidium. Each one of the four haploid nuclei migrates into its own basidiospore. The basidiospores are ballistically discharged and start new haploid mycelia called monokaryons. There are no males or females, rather there are compatible thalli with
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Many variations occur: some variations are self-compatible and spontaneously form dikaryons without a separate compatible thallus being involved. These fungi are said to be homothallic, versus the normal heterothallic species with mating types. Others are secondarily homothallic, in that two
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are examples of pathogenic basidiomycota. Such pathogens must be able to overcome the oxidative defenses of their respective hosts in order to produce a successful infection. The ability to undergo meiosis may provide a survival benefit for these fungi by promoting successful infection. A
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There are several genera classified in the Basidiomycota that are 1) poorly known, 2) have not been subjected to DNA analysis, or 3) if analysed phylogenetically do not group with as yet named or identified families, and have not been assigned to a specific family (i.e., they are
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Basidiomycota. In plant parasitic taxa, the saprotrophic phase is normally the yeast while the infectious stage is hyphal. However, there are examples of animal and human parasites where the species are dimorphic but it is the yeast-like state that is infectious. The genus
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because meiosis progresses synchronously in about 10 million cells within the mushroom cap, and the meiotic prophase stage is prolonged. Burns et al. studied the expression of genes involved in the 15-hour meiotic process, and found that the pattern of gene expression of
1303:), and haploid yeast-like conidia/basidiospores sometimes called sporidia, bud off laterally from each cell. In various smuts, the yeast phase may proliferate, or they may fuse, or they may infect plant tissue and become hyphal. In other smuts, such as 1250:
in dry chains in inverted cup-shaped bodies embedded in host tissue. These aeciospores then infect the second host, known as the primary or asexual host (in macrocyclic rusts). On the primary host a repeating spore stage is formed, numbered "II", the
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such as rain carry the spermatia from spermagonium to spermagonium, cross inoculating the mating types. Neither thallus is male or female. Once crossed, the dikaryons are established and a second spore stage is formed, numbered "I" and called
1213:) at their greatest complexity, produce five different types of spores on two different host plants in two unrelated host families. Such rusts are heteroecious (requiring two hosts) and macrocyclic (producing all five spores types). Wheat 1605:
Hibbett, David S.; Binder, Manfred; Bischoff, Joseph F.; Blackwell, Meredith; Cannon, Paul F.; Eriksson, Ove E.; Huhndorf, Sabine; James, Timothy; Kirk, Paul M. (May 2007). "A higher-level phylogenetic classification of the Fungi".
1291:. The teliospores are initially dikaryotic but become diploid via karyogamy. Meiosis takes place at the time of germination. A promycelium is formed that consists of a short hypha (equated to a basidium). In some smuts such as 1225:, which are miniature, flask-shaped, hollow, submicroscopic bodies embedded in the host tissue (such as a leaf). This stage, numbered "0", produces single-celled spores that ooze out in a sweet liquid and that act as nonmotile 1099:. These similarities in the patterns of expression led to the conclusion that the core expression program of meiosis has been conserved in these fungi for over half a billion years of evolution since these species diverged. 1032:. In most species, the basidiospores disperse and each can start a new haploid mycelium, continuing the lifecycle. Basidia are microscopic but they are often produced on or in multicelled large fructifications called 330:
reproducers. Basidiomycota that reproduce asexually (discussed below) can typically be recognized as members of this division by gross similarity to others, by the formation of a distinctive anatomical feature (the
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which are by then dikaryotic. Dikaryotic conidia may then form. Eventually the host is infected by infectious hyphae. Teliospores form in host tissue. Many variations on these general themes occur.
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Burns, C.; Stajich, J. E.; Rechtsteiner, A.; Casselton, L.; Hanlon, S. E.; Wilke, S. K.; Savytskyy, O. P.; Gathman, A. C.; Lilly, W. W.; Lieb, J. D.; Zolan, M. E.; Pukkila, P. J. (September 2010).
1356:, which are the asexual stages. Some Basidiomycota are only known as anamorphs. Many are yeasts, collectively called basidiomycetous yeasts to differentiate them from ascomycetous yeasts in the 1727: 1259:. Urediospores are dikaryotic and can infect the same host that produced them. They repeatedly infect this host over the growing season. At the end of the season, a fourth spore type, the 1287:
is the thick-walled, often darkly pigmented, ornate, teliospore that serves to survive harsh conditions such as overwintering and also serves to help disperse the fungus as dry
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follows shortly with the production of 4 haploid nuclei that migrate into 4 external, usually apical basidiospores. Variations occur, however. Typically the basidiospores are
575:, which includes mirror yeasts), and a mixture of odd, infrequently seen, or seldom recognized fungi, often parasitic on plants. The eight classes in the Pucciniomycotina are 530:. This sub-phyla also includes the "classic" mushrooms, polypores, corals, chanterelles, crusts, puffballs and stinkhorns. The three classes in the Agaricomycotina are the 1008:
nuclear divisions. Variations are frequent and multiple. In a typical Basidiomycota lifecycle the long lasting dikaryons periodically (seasonally or occasionally) produce
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Li, Liande; Wright, Sara J.; Krystofova, Svetlana; Park, Gyungsoon; Borkovich, Katherine A. (2007). "Heterotrimeric G Protein Signaling in Filamentous Fungi".
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characteristic central feature of meiosis is recombination between homologous chromosomes. This process is associated with repair of DNA damage, particularly
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after meiosis, with each of the 4 cells bearing one basidiospore each. The basidiospores disperse and start the infection process on host 1 again.
1173:. Occasionally, basidiospores are not formed and parts of the "basidia" act as the dispersal agents, e.g. the peculiar mycoparasitic jelly fungus, 702: 3074: 3026: 662: 2189: 4490: 3215: 2723: 2391: 842: 822: 782: 762: 5666: 3069: 1673:
Wijayawardene, Nalin; Hyde, Kevin; Al-Ani, Laith Khalil Tawfeeq; Somayeh, Dolatabadi; Stadler, Marc; Haelewaters, Danny; et al. (2020).
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the nuclei migrate into the promycelium that becomes septate (i.e., divided into cellular compartments separated by cell walls called
872: 692: 2142:"Light microscopy of basidia, basidiospores, and nuclei in spores and hyphae of Filobasidiella neoformans (Cryptococcus neoformans)" 5614: 1460:
Moore, R. T. (1980). "Taxonomic proposals for the classification of marine yeasts and other yeast-like fungi including the smuts".
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Other variations occur: some as standard lifecycles (that themselves have variations within variations) within specific orders.
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to undergo meiosis may contribute to their virulence by repairing the oxidative DNA damage caused by their host's release of
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given locus, and in such species, depending on the specifics, over 90% of monokaryons could be compatible with each other.
3438: 1309:, the elongated haploid basidiospores form apically, often in compatible pairs that fuse centrally resulting in H-shaped 1766:"Genome and transcriptome sequencing of the halophilic fungus Wallemia ichthyophaga: haloadaptations present and absent" 2384: 1943:; Heitman, J. (April 2005). "Sexual reproduction between partners of the same mating type in Cryptococcus neoformans". 932:
Unlike animals and plants which have readily recognizable male and female counterparts, Basidiomycota (except for the
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recognized three subphyla (Pucciniomycotina, Ustilaginomycotina, Agaricomycotina) and two other class level taxa (
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rusts complete their life-cycles on one host instead of two, and microcyclic rusts cut out one or more stages.
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The maintenance of the dikaryotic status in dikaryons in many Basidiomycota is facilitated by the formation of
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Scheme of a typical basidiocarp, the dipoid reproductive structure of a basidiomycete, showing fruiting body,
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that physically appear to help coordinate and re-establish pairs of compatible nuclei following synchronous
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Rodriguez-Carres, Marianela; Findley, Keisha; Sun, Sheng; Dietrich, Fred S.; Heitman, Joseph (2010-03-10).
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Michod, R. E.; Bernstein, H.; Nedelcu, A. M. (May 2008). "Adaptive value of sex in microbial pathogens".
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and then the compatible nuclei migrate into each other's mycelia and pair up with the resident nuclei.
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is not yet placed in a subdivision, but recent genomic evidence suggests that it is a sister group of
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The dimorphic Basidiomycota with yeast stages and the pleiomorphic rusts are examples of fungi with
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species can have two-, four-, six-, or eight-spored basidia, and the cultivated button mushroom,
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Zajc, J.; Liu, Y.; Dai, W.; Yang, Z.; Hu, J.; Gostinčar, C.; Gunde-Cimerman, N. (Sep 13, 2013).
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Prillinger, Hansjörg; Lopandic, Ksenija; Schweigkofler, Wolfgang; et al. (February 2002).
1617: 1317: 5771: 5702: 5178: 4828: 3519: 3036: 2698: 2667: 2479: 2408: 2292: 1333: 1238: 1187: 484: 291: 55: 5694: 1012:, the specialized usually club-shaped end cells, in which a pair of compatible nuclei fuse ( 5717: 5575: 4843: 3324: 3271: 2830: 2693: 2317: 2094: 1954: 1608: 1529: 1415: 576: 459: 372: 327: 8: 4334: 4118: 3301: 3276: 3016: 2817: 2756: 2751: 1345: 1181: 1115: 1074: 592: 424: 392: 214: 4706: 2098: 1958: 964:. The hyphae are then said to be dikaryotic. Conversely, the haploid mycelia are called 5173: 4855: 4344: 3536: 3514: 3502: 3477: 3424: 3377: 3329: 3316: 3296: 3281: 3261: 3210: 2881: 2853: 2807: 2743: 2496: 2474: 2462: 2437: 2117: 2082: 1978: 1916: 1887: 1794: 1765: 1643: 1580: 1293: 1109: 616: 596: 580: 572: 523: 515: 490: 432: 416: 408: 380: 376: 141: 63: 5554: 2166: 2141: 5781: 5689: 5562: 4965: 4678: 4604: 4585: 4295: 3878: 3546: 3509: 3497: 3487: 3344: 3339: 3334: 3286: 3137: 2987: 2873: 2799: 2506: 2469: 2457: 2447: 2208: 2171: 2122: 2054: 2015: 1970: 1940: 1921: 1841: 1799: 1635: 1560: 1541: 1533: 1409: 1362: 1288: 1153: 1037: 628: 624: 584: 436: 404: 400: 384: 364: 2034: 1691: 1674: 1584: 1327:
forms basidia on hyphae but the main infectious stage is more commonly known by the
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Traditionally, the Basidiomycota were divided into two classes, now obsolete:
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Smuts with both a yeast phase and an infectious hyphal state are examples of
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is an example. By convention, the stages and spore states are numbered by
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or the entire "basidium" acts as a "spore", e.g. in some false puffballs (
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is a basidiomycete mushroom. It is particularly suited to the study of
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Nonetheless these former concepts continue to be used as two types of
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is delayed, so that the compatible nuclei remain in pairs, called a
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include the rust fungi, the insect parasitic/symbiotic genus
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released via dissolution or disintegration of the basidia.
205: 199: 190: 184: 178: 163: 453:(alternatively called holobasidiomycetes), including true 237:. More specifically, Basidiomycota includes these groups: 4736: 988:) mating system. This results in the fact that following 172: 1816: 1666: 619:
are most (but not all) of the former smut fungi and the
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Kirk, P. M.; Cannon, P. F.; Stalpers, J. A. (2008).
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often has six-spored basidia, while some corticioid
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was similar to two other fungal species, the yeasts
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with respect to familial placement). These include:
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A 2007 classification, adopted by a coalition of 67
202: 175: 160: 952:. Typically haploid Basidiomycota mycelia fuse via 193: 187: 2340: 1870: 1858: 1751: 1660: 2258: 2256: 1559:Rivera-Mariani, F.E.; Bolaños-Rosero, B. (2011). 5758: 5023:Archaeal Richmond Mine acidophilic nanoorganisms 1407:infectious, state and the coffee leaf parasite, 623:. The classes of the Ustilaginomycotina are the 298:Basidiomycota are filamentous fungi composed of 4595: 2026: 1881: 1879: 1505: 1503: 2623: 2253: 2076: 2074: 1938: 1763: 4722: 3432: 2385: 1283:The characteristic part of the life-cycle of 1044:, etc. Ballistic basidiospores are formed on 3446: 1989: 1932: 1876: 1500: 506:include what had previously been called the 4311: 2264:"LUXGENE.COM: The glow-in-the-dark website" 2071: 1654: 1133: 912:Sexual reproduction cycle of basidiomycetes 326:. However, some Basidiomycota are obligate 181: 4729: 4715: 3439: 3425: 2392: 2378: 40: 3689: 2165: 2133: 2116: 2106: 1915: 1905: 1793: 1783: 1700: 1690: 1621: 727:Ammirati, A.D.Parker & Matheny (2007) 3594: 1366:with its apple-seed-shaped and coloured 1051: 928:Cell cycle of a Dikaryotic basidiomycete 923: 915: 907: 697:Bernicchia, Gorjón & Nakasone (2011) 4754: 3579: 1675:"Outline of Fungi and fungus-like taxa" 1028:, hence they are sometimes also called 777:Tkalčec, Mešić & Chun Y.Deng (2015) 571:, a former group of smut fungi (in the 14: 5759: 4982: 4697:denotes groups with over 1,000 species 2213:: CS1 maint: archived copy as title ( 1598: 1530:10.1146/annurev.micro.61.080706.093432 5501: 5500: 4710: 3420: 2373: 1459: 903: 610: 5682:05d71028-e72e-49e8-ad74-0211ea111628 2139: 1453: 972:male nor female. They have either a 1717:"Lecture 7 : Ustilaginomycets" 657:W.P.Wu, B.Sutton & Gange (1997) 556: 443:) with multiple orders and genera. 24: 634: 497: 318:(usually four). These specialized 25: 5803: 2350: 1999:Infection, Genetics and Evolution 1185:). In the human pathogenic genus 627:, the Entorrhizomycetes, and the 482:groupings, the "mushrooms" (e.g. 350: 1871:Kirk, Cannon & Stalpers 2008 1859:Kirk, Cannon & Stalpers 2008 1752:Kirk, Cannon & Stalpers 2008 1661:Kirk, Cannon & Stalpers 2008 339:components, and definitively by 156: 67: 2366:at the Tree of Life Web Project 2306: 2281: 2221: 2182: 1864: 1852: 1810: 1757: 1733:from the original on 2020-07-30 847:Hjortstam & Ryvarden (2008) 707:Hjortstam & Ryvarden (2008) 4176:      4172:      3932:      3928:      1745: 1709: 1552: 1480: 1437:List of Basidiomycota families 1040:, variously called mushrooms, 948:being composed of filamentous 488:) and the non-mushrooms (e.g. 295:, the human pathogenic yeast. 13: 1: 5787:Taxa named by Royall T. Moore 2158:10.1128/jb.128.1.445-455.1976 1513:Annual Review of Microbiology 1447: 767:Dhingra & Priyanka (2011) 2140:Erke, K. H. (October 1976). 2108:10.1371/journal.pone.0009620 2012:10.1016/j.meegid.2008.01.002 1907:10.1371/journal.pgen.1001135 1632:10.1016/j.mycres.2007.03.004 1376:with its sclerotium and its 817:Marvanová & Bärl. (2000) 687:Marvanová & Bärl. (1998) 667:Marvanová & Bärl. (2000) 314:that normally bear external 221:, constitute the subkingdom 7: 1825:Fungal Genetics and Biology 1425: 225:(often referred to as the " 10: 5808: 2334: 1068: 797:K.Ando & Tubaki (1984) 526:), as well as most of the 5509: 5472: 5290: 5207: 5159: 5044: 5035: 4978: 4750: 4746: 4741:phyla/divisions by domain 4673: 4641: 4618: 4545: 4514: 4474: 4445: 4425: 4416: 4404: 4400: 4245: 4236: 4207: 4192: 4183: 4170: 4166: 4099: 4071: 4056: 4047: 4006: 3985: 3976: 3939: 3926: 3922: 3909: 3896: 3892: 3869: 3849: 3845: 3841: 3787: 3774: 3765: 3732: 3719: 3685: 3674: 3656: 3643: 3590: 3575: 3571: 3455: 3403: 3353: 3315: 3252: 3237: 3186: 3128: 3107: 3098: 3050: 3007: 2996: 2978: 2969: 2931: 2908: 2890: 2872: 2839: 2816: 2798: 2780: 2769: 2742: 2707: 2684: 2666: 2654: 2645: 2616: 2582: 2565: 2556: 2538: 2531: 2415: 1838:10.1016/j.fgb.2012.01.007 1692:10.5943/mycosphere/11/1/8 1577:10.1007/s10453-011-9234-y 1493:Dictionary.com Unabridged 1096:Schizosaccharomyces pombe 128: 123: 64:Scientific classification 62: 48: 39: 34: 3902:     3898:     1785:10.1186/1471-2164-14-617 1379:Tilachlidiopsis racemosa 1278: 1246:, which form dikaryotic 1197: 1134:Variations in lifecycles 1090:Saccharomyces cerevisiae 992:, the resulting haploid 920:Basidiomycota life cycle 217:that, together with the 5777:Fungi by classification 4951:Thermodesulfobacteriota 3292:Cryptomycocolacomycetes 3085:Sporopachydermiomycetes 2900:Calcarisporiellomycetes 2343:Dictionary of the Fungi 2146:Journal of Bacteriology 1474:10.1515/bot-1980-230605 1373:Dendrocollybia racemosa 1340:Cryptococcus neoformans 1255:in dry pustules called 1128:reactive oxygen species 1104:Cryptococcus neoformans 589:Cryptomycocolacomycetes 389:Cryptomycocolacomycetes 233:. Members are known as 5792:Taxa described in 1980 2942:Palaeovesiarbusmycetes 2892:Calcarisporiellomycota 2859:Ramicandelaberomycetes 2676:Neocallimastigomycetes 2345:(10th ed.). CABI. 1229:, and also protruding 1209:, previously known as 1061: 929: 921: 913: 343:molecular analysis of 229:") within the kingdom 213:) is one of two large 5703:Paleobiology Database 5179:Neocallimastigomycota 3409:(polyphyletic group). 3037:Schizosaccharomycetes 2699:Monoblepharidomycetes 2668:Neocallimastigomycota 1724:Cpb-us-e1.wpmucdn.com 1055: 927: 919: 911: 485:Schizophyllum commune 124:Subdivisions/Classes 56:Kunstformen der Natur 4844:Coprothermobacterota 3325:Peribolosporomycetes 3272:Agaricostilbomycetes 3158:Collemopsidiomycetes 3032:Pneumocystidomycetes 2831:Entomophthoromycetes 2694:Hyaloraphidiomycetes 2575:Paramicrosporidiales 2241:on 27 September 2007 1609:Mycological Research 1416:Decapitatus flavidus 1116:double-strand breaks 577:Agaricostilbomycetes 460:Heterobasidiomycetes 373:Agaricostilbomycetes 49:Basidiomycetes from 4574:Gunflint microbiota 4335:Klebsormidiophyceae 4119:Hyphochytriomycetes 3302:Spiculogloeomycetes 3277:Cystobasidiomycetes 3090:Trigonopsidomycetes 3060:Alloascoideomycetes 3017:Archaeorhizomycetes 2952:Archaeosporomycetes 2818:Entomophthoromycota 2757:Blastocladiomycetes 2752:Physodermatomycetes 2718:Caulochytriomycetes 2558:Microsporidiomycota 2099:2010PLoSO...5.9620R 2039:Chemical Immunology 1967:10.1038/nature03448 1959:2005Natur.434.1017L 1346:Cryptococcus gattii 1237:and probably other 1075:Coprinopsis cinerea 593:Cystobasidiomycetes 425:Spiculogloeomycetes 393:Cystobasidiomycetes 304:basidiomycota-yeast 5174:Blastocladiomycota 4345:Coleochaetophyceae 3725:    3721:    3649:    3645:    3378:Bartheletiomycetes 3330:Monilielliomycetes 3317:Ustilaginomycotina 3297:Atractiellomycetes 3282:Microbotryomycetes 3262:Tritirachiomycetes 3211:Laboulbeniomycetes 2947:Paraglomeromycetes 2882:Mortierellomycetes 2854:Dimargaritomycetes 2808:Basidiobolomycetes 2744:Blastocladiomycota 2686:Monoblepharomycota 2362:2020-11-26 at the 1873:, pp. 717–718 1294:Mycosarcoma maydis 1118:. The ability of 1110:Mycosarcoma maydis 1062: 1036:or basidiomes, or 944:which are usually 930: 922: 914: 904:Typical life cycle 837:Ghob.-Nejh. (2010) 673:Anthoseptobasidium 617:Ustilaginomycotina 611:Ustilaginomycotina 597:Microbotryomycetes 581:Atractiellomycetes 573:Microbotryomycetes 491:Mycosarcoma maydis 451:Homobasidiomycetes 433:Tritirachiomycetes 417:Monilielliomycetes 409:Microbotryomycetes 381:Bartheletiomycetes 377:Atractiellomycetes 142:Ustilaginomycotina 5754: 5753: 5690:Open Tree of Life 5503:Taxon identifiers 5494: 5493: 5490: 5489: 5468: 5467: 5031: 5030: 4974: 4973: 4966:Verrucomicrobiota 4704: 4703: 4669: 4668: 4605:Spironematellidae 4586:Meteora sporadica 4541: 4540: 4537: 4536: 4470: 4469: 4396: 4395: 4392: 4391: 4388: 4387: 4384: 4383: 4380: 4379: 4376: 4375: 4372: 4371: 4368: 4367: 4296:Prasinodermophyta 4232: 4231: 4162: 4161: 4158: 4157: 4154: 4153: 4150: 4149: 4146: 4145: 4043: 4042: 3879:Centroplasthelida 3837: 3836: 3833: 3832: 3829: 3828: 3825: 3824: 3821: 3820: 3817: 3816: 3813: 3812: 3715: 3714: 3639: 3638: 3414: 3413: 3399: 3398: 3395: 3394: 3391: 3390: 3345:Exobasidiomycetes 3340:Ustilaginomycetes 3335:Malasseziomycetes 3287:Classiculomycetes 3233: 3232: 3229: 3228: 3178:Xylobotryomycetes 3138:Candelariomycetes 2988:Entorrhizomycetes 2965: 2964: 2874:Mortierellomycota 2800:Basidiobolomycota 2765: 2764: 2729:Sanchytriomycetes 2641: 2640: 2612: 2611: 2608: 2607: 2051:10.1159/000058868 2045:(9413): 207–295. 1953:(7036): 1017–21. 1410:Mycena citricolor 1363:Collybia tuberosa 1154:Agaricus bisporus 1002:clamp connections 898: 888: 878: 868: 867:W.Y.Zhuang (2008) 858: 848: 838: 828: 818: 808: 798: 788: 778: 768: 758: 748: 738: 737:P. Roberts (2006) 728: 718: 708: 698: 688: 678: 668: 658: 629:Ustilaginomycetes 625:Exobasidiomycetes 585:Classiculomycetes 437:Ustilaginomycetes 405:Malasseziomycetes 401:Exobasidiomycetes 385:Classiculomycetes 365:Entorrhizomycetes 149: 148: 119: 27:Division of fungi 16:(Redirected from 5799: 5747: 5746: 5734: 5733: 5721: 5720: 5711: 5710: 5698: 5697: 5685: 5684: 5675: 5674: 5662: 5661: 5649: 5648: 5636: 5635: 5623: 5622: 5610: 5609: 5597: 5596: 5584: 5583: 5571: 5570: 5558: 5557: 5545: 5544: 5543: 5530: 5529: 5528: 5498: 5497: 5242:Anthocerotophyta 5121:Hemimastigophora 5042: 5041: 5013:Nitrososphaerota 4980: 4979: 4956:Thermomicrobiota 4896:Kiritimatiellota 4891:Ignavibacteriota 4856:Deferribacterota 4819:Campylobacterota 4804:Bdellovibrionota 4752: 4751: 4748: 4747: 4731: 4724: 4717: 4708: 4707: 4651:Collodictyonidae 4597:Hemimastigophora 4593: 4592: 4552: 4423: 4422: 4414: 4413: 4410: 4402: 4401: 4351:Zygnematophyceae 4309: 4308: 4289: 4273: 4272: 4243: 4242: 4190: 4189: 4181: 4180: 4177: 4168: 4167: 4054: 4053: 3983: 3982: 3937: 3936: 3933: 3924: 3923: 3907: 3906: 3903: 3894: 3893: 3867: 3866: 3854: 3847: 3846: 3843: 3842: 3797:Choanoflagellata 3772: 3771: 3730: 3729: 3726: 3687: 3686: 3683: 3682: 3679: 3654: 3653: 3650: 3592: 3591: 3588: 3587: 3584: 3577: 3576: 3573: 3572: 3484: 3464: 3441: 3434: 3427: 3418: 3417: 3407:fungi imperfecti 3254:Pucciniomycotina 3250: 3249: 3201:Geoglossomycetes 3143:Coniocybomycetes 3105: 3104: 3065:Dipodascomycetes 3052:Saccharomycotina 3009:Taphrinomycotina 3005: 3004: 2980:Entorrhizomycota 2976: 2975: 2864:Kickxellomycetes 2826:Neozygitomycetes 2778: 2777: 2734:Chytridiomycetes 2664: 2663: 2652: 2651: 2621: 2620: 2563: 2562: 2536: 2535: 2444: 2424: 2407:classification, 2394: 2387: 2380: 2371: 2370: 2346: 2329: 2328: 2326: 2325: 2316:. Archived from 2310: 2304: 2303: 2301: 2300: 2291:. Archived from 2285: 2279: 2278: 2276: 2275: 2266:. Archived from 2260: 2251: 2250: 2248: 2246: 2240: 2234:. Archived from 2233: 2225: 2219: 2218: 2212: 2204: 2202: 2201: 2192:. Archived from 2186: 2180: 2179: 2169: 2137: 2131: 2130: 2120: 2110: 2078: 2069: 2068: 2066: 2065: 2030: 2024: 2023: 1993: 1987: 1986: 1936: 1930: 1929: 1919: 1909: 1883: 1874: 1868: 1862: 1856: 1850: 1849: 1814: 1808: 1807: 1797: 1787: 1761: 1755: 1749: 1743: 1742: 1740: 1738: 1732: 1721: 1713: 1707: 1706: 1704: 1694: 1670: 1664: 1663:, pp. 78–79 1658: 1652: 1651: 1625: 1602: 1596: 1595: 1593: 1591: 1556: 1550: 1549: 1507: 1498: 1497: 1484: 1478: 1477: 1457: 1432:Forest pathology 1231:receptive hyphae 897:P.Roberts (1994) 896: 886: 876: 866: 856: 846: 836: 826: 816: 806: 796: 786: 776: 766: 756: 746: 743:Dacryomycetopsis 736: 726: 717:P.Roberts (2005) 716: 706: 696: 686: 676: 666: 656: 563:Pucciniomycotina 557:Pucciniomycotina 514:layers on their 462:, including the 333:clamp connection 212: 211: 208: 207: 204: 201: 198: 195: 192: 189: 186: 183: 180: 177: 174: 171: 168: 165: 162: 137:Pucciniomycotina 114: 72: 71: 44: 32: 31: 21: 5807: 5806: 5802: 5801: 5800: 5798: 5797: 5796: 5757: 5756: 5755: 5750: 5742: 5737: 5729: 5724: 5716: 5714: 5706: 5701: 5693: 5688: 5680: 5678: 5670: 5665: 5657: 5652: 5644: 5639: 5631: 5626: 5618: 5613: 5605: 5600: 5592: 5587: 5579: 5574: 5566: 5561: 5553: 5548: 5539: 5538: 5533: 5524: 5523: 5518: 5505: 5495: 5486: 5464: 5415:Gnathostomulida 5320:Xenacoelomorpha 5286: 5282:Flowering plant 5237:Marchantiophyta 5203: 5169:Chytridiomycota 5155: 5027: 4970: 4926:Planctomycetota 4886:Gemmatimonadota 4871:Elusimicrobiota 4764:Acidobacteriota 4742: 4735: 4705: 4700: 4665: 4637: 4628:Ancyromonadidae 4614: 4591: 4546: 4533: 4524:Malawimonadidae 4510: 4466: 4441: 4405: 4364: 4307: 4280: 4279: 4271: 4228: 4203: 4171: 4142: 4095: 4067: 4039: 4002: 3972: 3927: 3918: 3897: 3888: 3865: 3850: 3809: 3783: 3761: 3720: 3711: 3675: 3670: 3644: 3635: 3580: 3567: 3566: 3482: 3460: 3451: 3445: 3415: 3410: 3387: 3383:Tremellomycetes 3373:Wallemiomycetes 3355:Agaricomycotina 3349: 3311: 3307:Pucciniomycetes 3241: 3225: 3221:Sordariomycetes 3182: 3173:Lecanoromycetes 3163:Dothideomycetes 3153:Arthoniomycetes 3124: 3094: 3080:Saccharomycetes 3046: 3042:Taphrinomycetes 3022:Neolectomycetes 3000: 2992: 2961: 2927: 2918:Endogonomycetes 2904: 2886: 2868: 2841:Kickxellomycota 2835: 2812: 2794: 2773: 2761: 2738: 2709:Chytridiomycota 2703: 2680: 2659: 2637: 2625:Aphelidiomycota 2617:Aphelidiomyceta 2604: 2595:Metchnikovellea 2590:Chytridiopsidea 2578: 2569:Morellosporales 2552: 2527: 2526: 2442: 2420: 2411: 2398: 2364:Wayback Machine 2353: 2337: 2332: 2323: 2321: 2312: 2311: 2307: 2298: 2296: 2287: 2286: 2282: 2273: 2271: 2262: 2261: 2254: 2244: 2242: 2238: 2231: 2227: 2226: 2222: 2206: 2205: 2199: 2197: 2190:"Archived copy" 2188: 2187: 2183: 2138: 2134: 2079: 2072: 2063: 2061: 2031: 2027: 1994: 1990: 1937: 1933: 1900:(9): e1001135. 1884: 1877: 1869: 1865: 1857: 1853: 1815: 1811: 1762: 1758: 1750: 1746: 1736: 1734: 1730: 1719: 1715: 1714: 1710: 1671: 1667: 1659: 1655: 1623:10.1.1.626.9582 1603: 1599: 1589: 1587: 1557: 1553: 1508: 1501: 1488:"Basidiomycota" 1486: 1485: 1481: 1462:Botanica Marina 1458: 1454: 1450: 1442:Mating in fungi 1428: 1422:called gemmae. 1306:Tilletia caries 1281: 1200: 1176:Tetragoniomyces 1171:Physalacriaceae 1136: 1071: 906: 901: 833:Punctulariopsis 757:Vellinga (2011) 637: 635:Genera included 613: 605:Pucciniomycetes 559: 551:Agaricomycotina 547:Wallemiomycetes 540:Tremellomycetes 504:Agaricomycotina 500: 498:Agaricomycotina 441:Wallemiomycetes 429:Tremellomycetes 421:Pucciniomycetes 361:Wallemiomycetes 353: 159: 155: 132:Agaricomycotina 113: 66: 28: 23: 22: 15: 12: 11: 5: 5805: 5795: 5794: 5789: 5784: 5779: 5774: 5769: 5752: 5751: 5749: 5748: 5735: 5722: 5712: 5699: 5686: 5676: 5663: 5650: 5637: 5624: 5611: 5598: 5585: 5572: 5559: 5546: 5531: 5515: 5513: 5507: 5506: 5492: 5491: 5488: 5487: 5480: 5478: 5475:Incertae sedis 5470: 5469: 5466: 5465: 5463: 5462: 5457: 5452: 5447: 5442: 5437: 5432: 5427: 5422: 5420:Micrognathozoa 5417: 5412: 5410:Acanthocephala 5407: 5402: 5397: 5392: 5387: 5382: 5377: 5372: 5367: 5362: 5357: 5352: 5347: 5342: 5337: 5332: 5327: 5322: 5317: 5312: 5307: 5302: 5296: 5294: 5288: 5287: 5285: 5284: 5279: 5274: 5269: 5264: 5259: 5254: 5252:Lycopodiophyta 5249: 5244: 5239: 5234: 5229: 5224: 5219: 5213: 5211: 5205: 5204: 5202: 5201: 5196: 5191: 5186: 5181: 5176: 5171: 5165: 5163: 5157: 5156: 5154: 5153: 5148: 5143: 5138: 5133: 5128: 5123: 5118: 5113: 5108: 5103: 5098: 5093: 5088: 5083: 5078: 5073: 5068: 5066:Ancyromonadida 5063: 5058: 5052: 5050: 5039: 5033: 5032: 5029: 5028: 5026: 5025: 5020: 5018:Thermoproteota 5015: 5010: 5003: 4996: 4988: 4986: 4976: 4975: 4972: 4971: 4969: 4968: 4963: 4958: 4953: 4948: 4943: 4938: 4933: 4931:Pseudomonadota 4928: 4923: 4918: 4913: 4908: 4906:Mycoplasmatota 4903: 4901:Lentisphaerota 4898: 4893: 4888: 4883: 4881:Fusobacteriota 4878: 4876:Fibrobacterota 4873: 4868: 4863: 4858: 4853: 4846: 4841: 4836: 4831: 4826: 4821: 4816: 4811: 4806: 4801: 4796: 4791: 4786: 4781: 4779:Armatimonadota 4776: 4771: 4769:Actinomycetota 4766: 4760: 4758: 4744: 4743: 4734: 4733: 4726: 4719: 4711: 4702: 4701: 4699: 4698: 4692: 4685: 4681:excluded from 4674: 4671: 4670: 4667: 4666: 4664: 4663: 4658: 4653: 4647: 4645: 4639: 4638: 4636: 4635: 4633:Planomonadidae 4630: 4624: 4622: 4620:Ancyromonadida 4616: 4615: 4613: 4612: 4610:Paramastigidae 4607: 4601: 4599: 4590: 4589: 4582: 4576: 4570: 4562: 4555: 4553: 4549:Incertae sedis 4543: 4542: 4539: 4538: 4535: 4534: 4532: 4531: 4526: 4520: 4518: 4512: 4511: 4509: 4508: 4503: 4498: 4493: 4488: 4480: 4478: 4472: 4471: 4468: 4467: 4465: 4464: 4457: 4451: 4449: 4443: 4442: 4440: 4439: 4432: 4426: 4420: 4411: 4398: 4397: 4394: 4393: 4390: 4389: 4386: 4385: 4382: 4381: 4378: 4377: 4374: 4373: 4370: 4369: 4366: 4365: 4363: 4362: 4354: 4347: 4342: 4337: 4332: 4325: 4317: 4315: 4306: 4305: 4298: 4292: 4290: 4270: 4269: 4262: 4257: 4252: 4246: 4240: 4238:Archaeplastida 4234: 4233: 4230: 4229: 4227: 4226: 4221: 4213: 4211: 4205: 4204: 4202: 4201: 4193: 4187: 4178: 4164: 4163: 4160: 4159: 4156: 4155: 4152: 4151: 4148: 4147: 4144: 4143: 4141: 4140: 4135: 4128: 4121: 4116: 4111: 4105: 4103: 4097: 4096: 4094: 4093: 4088: 4083: 4077: 4075: 4069: 4068: 4066: 4065: 4057: 4051: 4045: 4044: 4041: 4040: 4038: 4037: 4032: 4029:Dinoflagellata 4025: 4023:Chrompodellida 4020: 4012: 4010: 4004: 4003: 4001: 4000: 3993: 3986: 3980: 3974: 3973: 3971: 3970: 3963: 3956: 3951: 3945: 3943: 3934: 3920: 3919: 3917: 3916: 3910: 3904: 3890: 3889: 3887: 3886: 3881: 3875: 3873: 3864: 3863: 3857: 3855: 3852:Diaphoretickes 3839: 3838: 3835: 3834: 3831: 3830: 3827: 3826: 3823: 3822: 3819: 3818: 3815: 3814: 3811: 3810: 3808: 3807: 3799: 3793: 3791: 3785: 3784: 3782: 3781: 3775: 3769: 3763: 3762: 3760: 3759: 3754: 3749: 3741: 3733: 3727: 3717: 3716: 3713: 3712: 3710: 3709: 3701: 3695: 3693: 3680: 3672: 3671: 3669: 3668: 3663: 3657: 3651: 3641: 3640: 3637: 3636: 3634: 3633: 3628: 3623: 3618: 3611: 3606: 3600: 3598: 3585: 3569: 3568: 3565: 3564: 3563: 3562: 3561: 3560: 3557:Mesomycetozoea 3554: 3549: 3539: 3529: 3528: 3527: 3522: 3517: 3512: 3507: 3506: 3505: 3493:Diaphoretickes 3490: 3485: 3480: 3475: 3470: 3465: 3457: 3456: 3453: 3452: 3450:classification 3444: 3443: 3436: 3429: 3421: 3412: 3411: 3404: 3401: 3400: 3397: 3396: 3393: 3392: 3389: 3388: 3386: 3385: 3380: 3375: 3370: 3368:Agaricomycetes 3365: 3363:Dacrymycetales 3359: 3357: 3351: 3350: 3348: 3347: 3342: 3337: 3332: 3327: 3321: 3319: 3313: 3312: 3310: 3309: 3304: 3299: 3294: 3289: 3284: 3279: 3274: 3269: 3264: 3258: 3256: 3247: 3235: 3234: 3231: 3230: 3227: 3226: 3224: 3223: 3218: 3213: 3208: 3203: 3198: 3192: 3190: 3188:Sordariomyceta 3184: 3183: 3181: 3180: 3175: 3170: 3168:Eurotiomycetes 3165: 3160: 3155: 3150: 3148:Lichinomycetes 3145: 3140: 3134: 3132: 3130:Dothideomyceta 3126: 3125: 3123: 3122: 3117: 3115:Orbiliomycetes 3111: 3109: 3102: 3100:Pezizomycotina 3096: 3095: 3093: 3092: 3087: 3082: 3077: 3072: 3067: 3062: 3056: 3054: 3048: 3047: 3045: 3044: 3039: 3034: 3029: 3024: 3019: 3013: 3011: 3002: 2994: 2993: 2991: 2990: 2984: 2982: 2973: 2967: 2966: 2963: 2962: 2960: 2959: 2957:Glomeromycetes 2954: 2949: 2944: 2937: 2935: 2929: 2928: 2926: 2925: 2920: 2914: 2912: 2906: 2905: 2903: 2902: 2896: 2894: 2888: 2887: 2885: 2884: 2878: 2876: 2870: 2869: 2867: 2866: 2861: 2856: 2851: 2849:Zoopagomycetes 2845: 2843: 2837: 2836: 2834: 2833: 2828: 2822: 2820: 2814: 2813: 2811: 2810: 2804: 2802: 2796: 2795: 2793: 2792: 2790:Olpidiomycetes 2786: 2784: 2775: 2774:(paraphyletic) 2767: 2766: 2763: 2762: 2760: 2759: 2754: 2748: 2746: 2740: 2739: 2737: 2736: 2731: 2726: 2720: 2713: 2711: 2705: 2704: 2702: 2701: 2696: 2690: 2688: 2682: 2681: 2679: 2678: 2672: 2670: 2661: 2660:(paraphyletic) 2649: 2643: 2642: 2639: 2638: 2636: 2635: 2629: 2627: 2618: 2614: 2613: 2610: 2609: 2606: 2605: 2603: 2602: 2597: 2592: 2586: 2584: 2580: 2579: 2577: 2576: 2573: 2572:Nucleophagales 2570: 2566: 2560: 2554: 2553: 2551: 2550: 2548:Rozellomycetes 2544: 2542: 2533: 2529: 2528: 2525: 2524: 2523: 2522: 2521: 2520: 2517:Mesomycetozoea 2514: 2509: 2499: 2489: 2488: 2487: 2482: 2477: 2472: 2467: 2466: 2465: 2453:Diaphoretickes 2450: 2445: 2440: 2435: 2430: 2425: 2417: 2416: 2413: 2412: 2397: 2396: 2389: 2382: 2374: 2368: 2367: 2352: 2351:External links 2349: 2348: 2347: 2336: 2333: 2331: 2330: 2305: 2280: 2252: 2220: 2181: 2152:(1): 445–455. 2132: 2070: 2025: 1988: 1931: 1875: 1863: 1851: 1809: 1756: 1744: 1708: 1665: 1653: 1616:(5): 509–547. 1597: 1551: 1522:Annual Reviews 1499: 1496:(Online). n.d. 1479: 1451: 1449: 1446: 1445: 1444: 1439: 1434: 1427: 1424: 1324:Filobasidiella 1280: 1277: 1219:Roman numerals 1199: 1196: 1169:, both in the 1135: 1132: 1070: 1067: 1030:ballistospores 987: 983: 979: 975: 971: 905: 902: 900: 899: 889: 879: 869: 859: 849: 839: 829: 819: 809: 799: 789: 779: 769: 759: 749: 739: 729: 719: 709: 703:Brevicellopsis 699: 689: 679: 669: 659: 648: 643:incertae sedis 636: 633: 612: 609: 558: 555: 532:Agaricomycetes 520:Gasteromycetes 499: 496: 476: 475: 457: 369:Agaricomycetes 352: 351:Classification 349: 235:basidiomycetes 147: 146: 145: 144: 139: 134: 126: 125: 121: 120: 108: 104: 103: 98: 94: 93: 88: 84: 83: 78: 74: 73: 60: 59: 46: 45: 37: 36: 35:Basidiomycota 26: 9: 6: 4: 3: 2: 5804: 5793: 5790: 5788: 5785: 5783: 5780: 5778: 5775: 5773: 5770: 5768: 5767:Basidiomycota 5765: 5764: 5762: 5745: 5740: 5736: 5732: 5727: 5723: 5719: 5718:basidiomycota 5713: 5709: 5704: 5700: 5696: 5691: 5687: 5683: 5677: 5673: 5668: 5664: 5660: 5655: 5651: 5647: 5642: 5638: 5634: 5629: 5625: 5621: 5616: 5612: 5608: 5603: 5599: 5595: 5590: 5586: 5582: 5577: 5573: 5569: 5564: 5560: 5556: 5551: 5547: 5542: 5541:Basidiomycota 5536: 5532: 5527: 5521: 5517: 5516: 5514: 5512: 5511:Basidiomycota 5508: 5504: 5499: 5485: 5484: 5479: 5477: 5476: 5471: 5461: 5458: 5456: 5453: 5451: 5448: 5446: 5443: 5441: 5438: 5436: 5433: 5431: 5428: 5426: 5423: 5421: 5418: 5416: 5413: 5411: 5408: 5406: 5403: 5401: 5398: 5396: 5393: 5391: 5388: 5386: 5383: 5381: 5378: 5376: 5373: 5371: 5368: 5366: 5363: 5361: 5358: 5356: 5353: 5351: 5348: 5346: 5343: 5341: 5338: 5336: 5333: 5331: 5328: 5326: 5323: 5321: 5318: 5316: 5313: 5311: 5308: 5306: 5303: 5301: 5298: 5297: 5295: 5293: 5289: 5283: 5280: 5278: 5275: 5273: 5270: 5268: 5265: 5263: 5260: 5258: 5255: 5253: 5250: 5248: 5245: 5243: 5240: 5238: 5235: 5233: 5230: 5228: 5225: 5223: 5220: 5218: 5215: 5214: 5212: 5210: 5206: 5200: 5199:Basidiomycota 5197: 5195: 5192: 5190: 5187: 5185: 5184:Glomeromycota 5182: 5180: 5177: 5175: 5172: 5170: 5167: 5166: 5164: 5162: 5158: 5152: 5149: 5147: 5146:Stramenopiles 5144: 5142: 5139: 5137: 5134: 5132: 5129: 5127: 5124: 5122: 5119: 5117: 5114: 5112: 5109: 5107: 5104: 5102: 5099: 5097: 5094: 5092: 5089: 5087: 5084: 5082: 5079: 5077: 5074: 5072: 5071:Apusomonadida 5069: 5067: 5064: 5062: 5059: 5057: 5054: 5053: 5051: 5048: 5043: 5040: 5038: 5034: 5024: 5021: 5019: 5016: 5014: 5011: 5008: 5007:Nanoarchaeota 5004: 5001: 4997: 4994: 4993:Euryarchaeota 4990: 4989: 4987: 4985: 4981: 4977: 4967: 4964: 4962: 4959: 4957: 4954: 4952: 4949: 4947: 4944: 4942: 4941:Spirochaetota 4939: 4937: 4936:Rhodothermota 4934: 4932: 4929: 4927: 4924: 4922: 4919: 4917: 4914: 4912: 4909: 4907: 4904: 4902: 4899: 4897: 4894: 4892: 4889: 4887: 4884: 4882: 4879: 4877: 4874: 4872: 4869: 4867: 4866:Dictyoglomota 4864: 4862: 4859: 4857: 4854: 4851: 4850:Cyanobacteria 4847: 4845: 4842: 4840: 4839:Chrysiogenota 4837: 4835: 4834:Chloroflexota 4832: 4830: 4827: 4825: 4822: 4820: 4817: 4815: 4814:Calditrichota 4812: 4810: 4809:Caldisericota 4807: 4805: 4802: 4800: 4797: 4795: 4792: 4790: 4787: 4785: 4784:Atribacterota 4782: 4780: 4777: 4775: 4772: 4770: 4767: 4765: 4762: 4761: 4759: 4757: 4753: 4749: 4745: 4740: 4732: 4727: 4725: 4720: 4718: 4713: 4712: 4709: 4696: 4693: 4690: 4686: 4684: 4680: 4677:¹traditional 4676: 4675: 4672: 4662: 4659: 4657: 4656:Mantamonadida 4654: 4652: 4649: 4648: 4646: 4644: 4640: 4634: 4631: 4629: 4626: 4625: 4623: 4621: 4617: 4611: 4608: 4606: 4603: 4602: 4600: 4598: 4594: 4588: 4587: 4583: 4581: 4577: 4575: 4571: 4569: 4568: 4563: 4561: 4557: 4556: 4554: 4551: 4550: 4544: 4530: 4527: 4525: 4522: 4521: 4519: 4517: 4513: 4507: 4504: 4502: 4499: 4497: 4494: 4492: 4489: 4487: 4486: 4482: 4481: 4479: 4477: 4473: 4463: 4462: 4458: 4456: 4455:Heterolobosea 4453: 4452: 4450: 4448: 4447:Discicristata 4444: 4438: 4437: 4433: 4431: 4428: 4427: 4424: 4421: 4419: 4415: 4412: 4408: 4403: 4399: 4360: 4359: 4355: 4353: 4352: 4348: 4346: 4343: 4341: 4338: 4336: 4333: 4331: 4330: 4326: 4324: 4323: 4319: 4318: 4316: 4314: 4310: 4304: 4303: 4299: 4297: 4294: 4293: 4291: 4287: 4286: 4278: 4277:Viridiplantae 4274: 4268: 4267: 4263: 4261: 4258: 4256: 4253: 4251: 4248: 4247: 4244: 4241: 4239: 4235: 4225: 4222: 4220: 4219: 4215: 4214: 4212: 4210: 4206: 4200: 4199: 4198:Microheliella 4195: 4194: 4191: 4188: 4186: 4182: 4179: 4175: 4169: 4165: 4139: 4138:Actinophryida 4136: 4134: 4133: 4129: 4127: 4126: 4122: 4120: 4117: 4115: 4112: 4110: 4107: 4106: 4104: 4102: 4098: 4092: 4089: 4087: 4084: 4082: 4079: 4078: 4076: 4074: 4070: 4064: 4063: 4059: 4058: 4055: 4052: 4050: 4049:Stramenopiles 4046: 4036: 4033: 4031: 4030: 4026: 4024: 4021: 4019: 4018: 4014: 4013: 4011: 4009: 4005: 3999: 3998: 3994: 3991: 3988: 3987: 3984: 3981: 3979: 3975: 3969: 3968: 3964: 3962: 3961: 3957: 3955: 3952: 3950: 3947: 3946: 3944: 3942: 3938: 3935: 3931: 3925: 3921: 3915: 3912: 3911: 3908: 3905: 3901: 3895: 3891: 3885: 3882: 3880: 3877: 3876: 3874: 3872: 3868: 3862: 3859: 3858: 3856: 3853: 3848: 3844: 3840: 3805: 3804: 3800: 3798: 3795: 3794: 3792: 3790: 3786: 3780: 3777: 3776: 3773: 3770: 3768: 3764: 3758: 3755: 3753: 3752:Ichthyosporea 3750: 3748: 3747: 3742: 3740: 3739: 3735: 3734: 3731: 3728: 3724: 3718: 3707: 3706: 3702: 3700: 3699:Rotosphaerida 3697: 3696: 3694: 3692: 3688: 3684: 3681: 3678: 3673: 3667: 3664: 3662: 3661:Apusomonadida 3659: 3658: 3655: 3652: 3648: 3642: 3632: 3629: 3627: 3624: 3622: 3619: 3617: 3616: 3612: 3610: 3607: 3605: 3602: 3601: 3599: 3597: 3593: 3589: 3586: 3583: 3578: 3574: 3570: 3558: 3555: 3553: 3550: 3548: 3545: 3544: 3543: 3540: 3538: 3535: 3534: 3533: 3530: 3526: 3523: 3521: 3520:Stramenopiles 3518: 3516: 3513: 3511: 3508: 3504: 3501: 3500: 3499: 3496: 3495: 3494: 3491: 3489: 3486: 3483:(major groups 3481: 3479: 3476: 3474: 3471: 3469: 3466: 3463: 3459: 3458: 3454: 3449: 3442: 3437: 3435: 3430: 3428: 3423: 3422: 3419: 3408: 3402: 3384: 3381: 3379: 3376: 3374: 3371: 3369: 3366: 3364: 3361: 3360: 3358: 3356: 3352: 3346: 3343: 3341: 3338: 3336: 3333: 3331: 3328: 3326: 3323: 3322: 3320: 3318: 3314: 3308: 3305: 3303: 3300: 3298: 3295: 3293: 3290: 3288: 3285: 3283: 3280: 3278: 3275: 3273: 3270: 3268: 3265: 3263: 3260: 3259: 3257: 3255: 3251: 3248: 3245: 3240: 3239:Basidiomycota 3236: 3222: 3219: 3217: 3214: 3212: 3209: 3207: 3206:Leotiomycetes 3204: 3202: 3199: 3197: 3196:Xylonomycetes 3194: 3193: 3191: 3189: 3185: 3179: 3176: 3174: 3171: 3169: 3166: 3164: 3161: 3159: 3156: 3154: 3151: 3149: 3146: 3144: 3141: 3139: 3136: 3135: 3133: 3131: 3127: 3121: 3120:Pezizomycetes 3118: 3116: 3113: 3112: 3110: 3106: 3103: 3101: 3097: 3091: 3088: 3086: 3083: 3081: 3078: 3076: 3075:Pichiomycetes 3073: 3071: 3068: 3066: 3063: 3061: 3058: 3057: 3055: 3053: 3049: 3043: 3040: 3038: 3035: 3033: 3030: 3028: 3027:Novakomycetes 3025: 3023: 3020: 3018: 3015: 3014: 3012: 3010: 3006: 3003: 2999: 2995: 2989: 2986: 2985: 2983: 2981: 2977: 2974: 2972: 2968: 2958: 2955: 2953: 2950: 2948: 2945: 2943: 2939: 2938: 2936: 2934: 2933:Glomeromycota 2930: 2924: 2923:Mucoromycetes 2921: 2919: 2916: 2915: 2913: 2911: 2907: 2901: 2898: 2897: 2895: 2893: 2889: 2883: 2880: 2879: 2877: 2875: 2871: 2865: 2862: 2860: 2857: 2855: 2852: 2850: 2847: 2846: 2844: 2842: 2838: 2832: 2829: 2827: 2824: 2823: 2821: 2819: 2815: 2809: 2806: 2805: 2803: 2801: 2797: 2791: 2788: 2787: 2785: 2783: 2782:Olpidiomycota 2779: 2776: 2772: 2768: 2758: 2755: 2753: 2750: 2749: 2747: 2745: 2741: 2735: 2732: 2730: 2727: 2725: 2721: 2719: 2715: 2714: 2712: 2710: 2706: 2700: 2697: 2695: 2692: 2691: 2689: 2687: 2683: 2677: 2674: 2673: 2671: 2669: 2665: 2662: 2657: 2653: 2650: 2648: 2644: 2634: 2631: 2630: 2628: 2626: 2622: 2619: 2615: 2601: 2598: 2596: 2593: 2591: 2588: 2587: 2585: 2583:Microsporidia 2581: 2574: 2571: 2568: 2567: 2564: 2561: 2559: 2555: 2549: 2546: 2545: 2543: 2541: 2540:Rozellomycota 2537: 2534: 2532:Rozellomyceta 2530: 2518: 2515: 2513: 2510: 2508: 2505: 2504: 2503: 2500: 2498: 2495: 2494: 2493: 2490: 2486: 2483: 2481: 2480:Stramenopiles 2478: 2476: 2473: 2471: 2468: 2464: 2461: 2460: 2459: 2456: 2455: 2454: 2451: 2449: 2446: 2443:(major groups 2441: 2439: 2436: 2434: 2431: 2429: 2426: 2423: 2419: 2418: 2414: 2410: 2409:fungal orders 2406: 2402: 2395: 2390: 2388: 2383: 2381: 2376: 2375: 2372: 2365: 2361: 2358: 2357:Basidiomycota 2355: 2354: 2344: 2339: 2338: 2320:on 2006-12-21 2319: 2315: 2309: 2295:on 2006-12-21 2294: 2290: 2284: 2270:on 2007-09-28 2269: 2265: 2259: 2257: 2237: 2230: 2224: 2216: 2210: 2196:on 2007-10-25 2195: 2191: 2185: 2177: 2173: 2168: 2163: 2159: 2155: 2151: 2147: 2143: 2136: 2128: 2124: 2119: 2114: 2109: 2104: 2100: 2096: 2092: 2088: 2084: 2077: 2075: 2060: 2056: 2052: 2048: 2044: 2040: 2036: 2029: 2021: 2017: 2013: 2009: 2006:(3): 267–85. 2005: 2001: 2000: 1992: 1984: 1980: 1976: 1972: 1968: 1964: 1960: 1956: 1952: 1948: 1947: 1942: 1935: 1927: 1923: 1918: 1913: 1908: 1903: 1899: 1895: 1894: 1893:PLOS Genetics 1889: 1882: 1880: 1872: 1867: 1861:, p. 581 1860: 1855: 1847: 1843: 1839: 1835: 1832:(3): 217–26. 1831: 1827: 1826: 1821: 1813: 1805: 1801: 1796: 1791: 1786: 1781: 1777: 1773: 1772: 1767: 1760: 1753: 1748: 1729: 1725: 1718: 1712: 1703: 1698: 1693: 1688: 1685:: 1060–1456. 1684: 1680: 1676: 1669: 1662: 1657: 1649: 1645: 1641: 1637: 1633: 1629: 1624: 1619: 1615: 1611: 1610: 1601: 1586: 1582: 1578: 1574: 1570: 1566: 1562: 1555: 1547: 1543: 1539: 1535: 1531: 1527: 1523: 1519: 1515: 1514: 1506: 1504: 1495: 1494: 1489: 1483: 1475: 1471: 1467: 1463: 1456: 1452: 1443: 1440: 1438: 1435: 1433: 1430: 1429: 1423: 1421: 1418: 1417: 1412: 1411: 1406: 1403: 1402: 1397: 1396: 1395:Hohenbuehelia 1391: 1387: 1386: 1381: 1380: 1375: 1374: 1369: 1365: 1364: 1359: 1355: 1350: 1348: 1347: 1342: 1341: 1336: 1335: 1330: 1326: 1325: 1319: 1314: 1312: 1308: 1307: 1302: 1301: 1296: 1295: 1290: 1286: 1276: 1274: 1270: 1266: 1262: 1258: 1254: 1249: 1245: 1240: 1236: 1232: 1228: 1224: 1220: 1216: 1212: 1208: 1204: 1195: 1192: 1190: 1189: 1184: 1183: 1178: 1177: 1172: 1168: 1167: 1162: 1161: 1156: 1155: 1150: 1149: 1144: 1143: 1131: 1129: 1125: 1121: 1120:C. neoformans 1117: 1112: 1111: 1106: 1105: 1100: 1098: 1097: 1092: 1091: 1086: 1081: 1077: 1076: 1066: 1059: 1054: 1050: 1047: 1043: 1039: 1035: 1031: 1027: 1023: 1019: 1015: 1011: 1007: 1003: 998: 995: 994:basidiospores 991: 985: 981: 977: 973: 969: 967: 963: 959: 955: 951: 947: 943: 939: 935: 926: 918: 910: 895: 894: 890: 885: 884: 880: 875: 874: 870: 865: 864: 860: 855: 854: 850: 845: 844: 840: 835: 834: 830: 827:Corner (1966) 825: 824: 820: 815: 814: 810: 807:Wakef. (1934) 805: 804: 800: 795: 794: 790: 787:Oberw. (1990) 785: 784: 780: 775: 774: 770: 765: 764: 760: 755: 754: 750: 745: 744: 740: 735: 734: 730: 725: 724: 720: 715: 714: 710: 705: 704: 700: 695: 694: 690: 685: 684: 680: 675: 674: 670: 665: 664: 663:Anguillomyces 660: 655: 654: 650: 649: 647: 645: 644: 632: 630: 626: 622: 621:Exobasidiales 618: 608: 606: 602: 598: 594: 590: 586: 582: 578: 574: 570: 569: 568:Septobasidium 564: 554: 552: 548: 543: 541: 537: 533: 529: 525: 521: 517: 513: 509: 508:Hymenomycetes 505: 495: 493: 492: 487: 486: 481: 473: 469: 465: 461: 458: 456: 452: 449: 448: 447: 444: 442: 438: 434: 430: 426: 422: 418: 414: 410: 406: 402: 398: 394: 390: 386: 382: 378: 374: 370: 366: 362: 358: 348: 346: 342: 338: 334: 329: 325: 324:basidiospores 321: 317: 313: 309: 305: 301: 296: 294: 293: 288: 287:mirror yeasts 284: 280: 276: 272: 268: 264: 260: 256: 252: 251:bracket fungi 248: 244: 240: 236: 232: 228: 224: 220: 216: 210: 153: 152:Basidiomycota 143: 140: 138: 135: 133: 130: 129: 127: 122: 117: 112: 111:Basidiomycota 109: 106: 105: 102: 99: 96: 95: 92: 89: 86: 85: 82: 79: 76: 75: 70: 65: 61: 58: 57: 52: 51:Ernst Haeckel 47: 43: 38: 33: 30: 19: 18:Basidiomycete 5772:Fungus phyla 5510: 5481: 5473: 5395:Orthonectida 5390:Gastrotricha 5365:Nematomorpha 5340:Chaetognatha 5330:Hemichordate 5257:Pteridophyta 5198: 5126:Malawimonada 5000:Korarchaeota 4961:Thermotogota 4946:Synergistota 4921:Nitrospirota 4916:Nitrospinota 4861:Deinococcota 4794:Bacteroidota 4694: 4689:paraphyletic 4584: 4565: 4547: 4516:Malawimonada 4496:Anaeromonada 4491:Anaeramoebae 4483: 4459: 4436:Tsukubamonas 4434: 4356: 4349: 4340:Charophyceae 4327: 4320: 4313:Streptophyta 4300: 4284: 4264: 4260:Rhodelphidia 4218:Palpitomonas 4216: 4196: 4185:Pancryptista 4130: 4123: 4060: 4027: 4015: 3995: 3965: 3960:Foraminifera 3958: 3801: 3744: 3738:Tunicaraptor 3736: 3703: 3677:Opisthokonta 3613: 3542:Opisthokonta 3267:Mixiomycetes 3238: 3216:Sareomycetes 2910:Mucoromycota 2724:Coenomycetes 2502:Opisthokonta 2342: 2322:. 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Retrieved 1571:(2): 83–97. 1568: 1565:Aerobiologia 1564: 1554: 1517: 1511: 1491: 1482: 1465: 1461: 1455: 1414: 1408: 1401:Nematoctonus 1399: 1393: 1383: 1377: 1371: 1361: 1351: 1344: 1338: 1334:Cryptococcus 1332: 1322: 1315: 1304: 1298: 1292: 1282: 1253:urediospores 1201: 1193: 1188:Cryptococcus 1186: 1180: 1174: 1164: 1158: 1152: 1146: 1140: 1137: 1123: 1119: 1108: 1102: 1101: 1094: 1088: 1084: 1073: 1072: 1063: 1060:and basidia. 1034:basidiocarps 1016:) to form a 999: 978:unifactorial 931: 891: 887:Corda (1837) 881: 871: 861: 857:Vánky (2008) 851: 843:Radulodontia 841: 831: 823:Paraphelaria 821: 811: 801: 791: 783:Kryptastrina 781: 771: 763:Hallenbergia 761: 751: 741: 731: 721: 711: 701: 691: 681: 671: 661: 651: 641: 638: 614: 601:Mixiomycetes 566: 560: 544: 536:Dacrymycetes 501: 489: 483: 480:growth habit 477: 445: 413:Mixiomycetes 397:Dacrymycetes 354: 345:DNA sequence 341:phylogenetic 302:(except for 297: 292:Cryptococcus 290: 267:chanterelles 234: 227:higher fungi 226: 151: 150: 110: 97:Subkingdom: 54: 29: 5628:iNaturalist 5535:Wikispecies 5450:Brachiopoda 5425:Cycliophora 5370:Onychophora 5345:Kinorhyncha 5267:Ginkgophyta 5262:Cycadophyta 5227:Chlorophyta 5217:Glaucophyta 4911:Myxococcota 4829:Chlorobiota 4824:Chlamydiota 4580:Chitinozoan 4506:Parabasalia 4358:Embryophyta 4322:Chlorokybus 4302:Chlorophyta 4250:Glaucophyta 4224:Cryptophyta 4114:Pirsoniales 4081:Bicosoecida 4062:Platysulcus 4035:Perkinsozoa 4017:Apicomplexa 3990:Colponemida 3757:Pluriformea 3626:Archamoebea 3615:Eumycetozoa 3070:Lipomycetes 3001:(sac fungi) 2600:Microsporea 2401:Opisthokont 1941:Hull, C. M. 1702:10481/61998 1524:: 423–452. 1398:with their 1390:rhizomorphs 1388:with their 1331:yeast name 1265:promycelium 1248:aeciospores 1207:Pucciniales 1182:Scleroderma 1142:Craterellus 1038:fruitbodies 986:bifactorial 966:monokaryons 938:Pucciniales 877:Rick (1958) 813:Nodulospora 793:Microstella 773:Hymenoporus 747:Rick (1958) 723:Cleistocybe 713:Celatogloea 677:Rick (1943) 653:Anastomyces 528:jelly fungi 524:fruitbodies 516:fruitbodies 357:mycologists 322:are called 271:earth stars 259:jelly fungi 5761:Categories 5483:Parakaryon 5445:Entoprocta 5380:Arthropoda 5375:Tardigrada 5355:Priapulida 5350:Loricifera 5335:Echinoderm 5305:Ctenophora 5277:Gnetophyta 5232:Charophyta 5222:Rhodophyta 5194:Ascomycota 5189:Zygomycota 5131:Metamonada 5106:Euglenozoa 5091:Ciliophora 4799:Balneolota 4774:Aquificota 4661:Rigifilida 4560:Acritarchs 4485:Barthelona 4476:Metamonada 4461:Euglenozoa 4329:Mesostigma 4285:sensu lato 4266:Rhodophyta 4125:Ochrophyta 4086:Placidozoa 3997:Ciliophora 3967:Radiolaria 3884:Haptophyta 3779:Filasterea 3691:Holomycota 3405:See also: 2998:Ascomycota 2771:Zygomycota 2405:True fungi 2324:2007-04-17 2299:2007-04-17 2289:"Hohenbue" 2274:2007-09-13 2200:2007-09-13 2064:2024-03-12 1679:Mycosphere 1590:16 January 1468:(6): 371. 1448:References 1420:propagules 1413:, and its 1385:Armillaria 1368:sclerotium 1358:Ascomycota 1329:anamorphic 1273:Autoecious 1261:teliospore 1211:Uredinales 1160:Armillaria 1148:Sistotrema 1085:C. cinerea 1046:sterigmata 982:tetrapolar 954:plasmogamy 883:Zygodesmus 803:Neotyphula 733:Cystogloea 545:The class 538:, and the 316:meiospores 247:stinkhorns 219:Ascomycota 107:Division: 5731:100364590 5550:AusLichen 5400:Dicyemida 5272:Pinophyta 5151:Telonemia 5086:Cryptista 5076:Breviatea 5061:Amoebozoa 5056:Alveolata 5037:Eukaryote 4789:Bacillota 4501:Fornicata 4407:Excavates 4209:Cryptista 4132:Oomycetes 4109:Developea 4091:Sagenista 3978:Alveolata 3914:Telonemia 3789:Choanozoa 3666:Breviatea 3609:Tubulinea 3596:Amoebozoa 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1269:septate 1239:vectors 1235:Insects 1080:meiosis 1069:Meiosis 1022:Meiosis 1018:diploid 1010:basidia 1006:mitotic 990:meiosis 980:) or a 974:bipolar 970:neither 946:mycelia 873:Zanchia 693:Arrasia 518:), the 328:asexual 312:basidia 310:called 263:boletes 239:agarics 223:Dikarya 101:Dikarya 5708:212608 5695:634628 5679:NZOR: 5646:623881 5594:1BASIP 5300:Sponge 5292:Animal 4691:groups 4073:Bigyra 3647:Obazoa 3525:Plants 3462:Domain 3242:(with 2485:Plants 2422:Domain 2176:789347 2174:  2167:232872 2164:  2125:  2115:  2057:  2018:  1981:  1973:  1946:Nature 1924:  1914:  1844:  1802:  1792:  1646:  1638:  1620:  1583:  1544:  1536:  1166:Xerula 1020:cell. 950:hyphae 603:, and 534:, the 347:data. 320:spores 300:hyphae 289:, and 118:(1980) 5744:21890 5739:WoRMS 5715:PPE: 5659:90050 5633:47169 5607:90050 5209:Plant 5161:Fungi 5081:CRuMs 4643:CRuMs 3705:Fungi 3552:Fungi 3108:Other 2658:fungi 2512:Fungi 2239:(PDF) 2232:(PDF) 1979:S2CID 1731:(PDF) 1720:(PDF) 1644:S2CID 1581:S2CID 1520:(1). 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Index

Basidiomycete

Ernst Haeckel
Kunstformen der Natur
Scientific classification
Edit this classification
Eukaryota
Fungi
Dikarya
Basidiomycota
R.T.Moore
Agaricomycotina
Pucciniomycotina
Ustilaginomycotina
/bəˌsɪdi.mˈktə/
divisions
Ascomycota
Dikarya
Fungi
agarics
puffballs
stinkhorns
bracket fungi
polypores
jelly fungi
boletes
chanterelles
earth stars
smuts
bunts

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