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Biome

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5686: 455: 5045: 5710: 1318: 5541: 2182: 267: 5674: 5698: 38: 202:(a concrete geographical unit), while the biome definition used in this article is used as an international, non-regional, terminology—irrespectively of the continent in which an area is present, it takes the same biome name—and corresponds to his "zonobiome", "orobiome" and "pedobiome" (biomes determined by climate zone, altitude or soil). 2160:
has the potential to greatly alter the distribution of Earth's biomes. Meaning, biomes around the world could change so much that they would be at risk of becoming new biomes entirely. More specifically, between 54% and 22% of global land area will experience climates that correspond to other biomes.
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Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Allnutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P., Kassem, K. R. (2001). Terrestrial
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for the United States in a map published in 1976. He subsequently expanded the system to include the rest of North America in 1981, and the world in 1989. The Bailey system, based on climate, is divided into four domains (polar, humid temperate, dry, and humid tropical), with further divisions based
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Whittaker summed the effects of gradients (3) and (4) to get an overall temperature gradient and combined this with a gradient (2), the moisture gradient, to express the above conclusions in what is known as the Whittaker classification scheme. The scheme graphs average annual precipitation (x-axis)
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To divide the world into a few ecological zones is difficult, notably because of the small-scale variations that exist everywhere on earth and because of the gradual changeover from one biome to the other. Their boundaries must therefore be drawn arbitrarily and their characterization made according
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Average temperatures have risen more than twice the usual amount in both arctic and mountainous biomes, which leads to the conclusion that arctic and mountainous biomes are currently the most vulnerable to climate change. South American terrestrial biomes have been predicted to go through the same
925:. The boundaries of each biome correlate to the conditions of moisture and cold stress that are strong determinants of plant form, and therefore the vegetation that defines the region. Extreme conditions, such as flooding in a swamp, can create different kinds of communities within the same biome. 3161:
Abell, R., M. Thieme, C. Revenga, M. Bryer, M. Kottelat, N. Bogutskaya, B. Coad, N. Mandrak, S. Contreras-Balderas, W. Bussing, M. L. J. Stiassny, P. Skelton, G. R. Allen, P. Unmack, A. Naseka, R. Ng, N. Sindorf, J. Robertson, E. Armijo, J. Higgins, T. J. Heibel, E. Wikramanayake, D. Olson, H. L.
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factors are the largest determinants of the types of vegetation found in a habitat. Holdridge uses the four axes to define 30 so-called "humidity provinces", which are clearly visible in his diagram. While this scheme largely ignores soil and sun exposure, Holdridge acknowledged that these were
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and temperature lead to below-ground primary production (roots), and temperature and water lead to cool and warm season growth habit. These findings help explain the categories used in Holdridge's bioclassification scheme (see below), which were then later simplified by Whittaker. The number of
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Nolan, Connor; Overpeck, Jonathan T.; Allen, Judy R. M.; Anderson, Patricia M.; Betancourt, Julio L.; Binney, Heather A.; Brewer, Simon; Bush, Mark B.; Chase, Brian M.; Cheddadi, Rachid; Djamali, Morteza; Dodson, John; Edwards, Mary E.; Gosling, William D.; Haberle, Simon (2018-08-31).
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A biota is the total collection of organisms of a geographic region or a time period, from local geographic scales and instantaneous temporal scales all the way up to whole-planet and whole-timescale spatiotemporal scales. The biotas of the Earth make up the
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Locosselli, Giuliano Maselli; Brienen, Roel J. W.; Leite, Melina de Souza; Gloor, Manuel; Krottenthaler, Stefan; Oliveira, Alexandre A. de; Barichivich, Jonathan; Anhuf, Dieter; Ceccantini, Gregorio; Schöngart, Jochen; Buckeridge, Marcos (2020-12-14).
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Martins, F. R. & Batalha, M. A. (2011). Formas de vida, espectro biológico de Raunkiaer e fisionomia da vegetação. In: Felfili, J. M., Eisenlohr, P. V.; Fiuza de Melo, M. M. R.; Andrade, L. A.; Meira Neto, J. A. A. (Org.).
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classification scheme considers the seasonality of temperature and precipitation. The system, also assessing precipitation and temperature, finds nine major biome types, with the important climate traits and
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classified biomes using two abiotic factors: precipitation and temperature. His scheme can be seen as a simplification of Holdridge's; more readily accessible, but missing Holdridge's greater specificity.
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processes. As a result, vegetation forms predicted by conventional biome systems can no longer be observed across much of Earth's land surface as they have been replaced by crop and rangelands or cities.
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Sims, Phillip L.; Singh, J.S. (July 1978). "The Structure and Function of Ten Western North American Grasslands: III. Net Primary Production, Turnover and Efficiencies of Energy Capture and Water Use".
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Changes in physiognomic complexity vary with how favorable of an environment exists (decreasing community structure and reduction of stratal differentiation as the environment becomes less favorable).
229:, a geographic space with subcontinental dimensions, with the predominance of similar geomorphologic and climatic characteristics, and of a certain vegetation form. Both include many biomes in fact. 258:
classification schemes and the variety of determinants used in those schemes, however, should be taken as strong indicators that biomes do not fit perfectly into the classification schemes created.
4172:"IPCC Special Report on Climate Change, Desertification, Land Degradation, Sustainable Land Management, Food Security, and Greenhouse gas fluxes in Terrestrial Ecosystems:Summary for Policymakers" 484:
Biome: a grouping of terrestrial ecosystems on a given continent that is similar in vegetation structure, physiognomy, features of the environment and characteristics of their animal communities.
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Dobrowski, Solomon Z.; Littlefield, Caitlin E.; Lyons, Drew S.; Hollenberg, Clark; Carroll, Carlos; Parks, Sean A.; Abatzoglou, John T.; Hegewisch, Katherine; Gage, Josh (September 29, 2021).
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Lopez, R. E. d. Reis, J. G. Lundberg, M. H. Sabaj Perez, and P. Petry. (2008). Freshwater ecoregions of the world: A new map of biogeographic units for freshwater biodiversity conservation.
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Holdridge life zone classification scheme. Although conceived as three-dimensional by its originator, it is usually shown as a two-dimensional array of hexagons in a triangular frame.
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only, while biome is used when concerned with both plants and animals. Whittaker's convention of biome-type or formation-type is a broader method to categorize similar communities.
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in mm/yr and above-ground net primary production in g/m/yr. The general results from the study were that precipitation and water use led to above-ground primary production, while
2091:. Anthropogenic biomes offer a way to recognize the irreversible coupling of human and ecological systems at global scales and manage Earth's biosphere and anthropogenic biomes. 481:: sometimes referring to the plants' appearance; or the biome's apparent characteristics, outward features, or appearance of ecological communities or species - including plants. 2668: 569: 4571: 3330: 2169:
temperature trends as arctic and mountainous biomes. With its annual average temperature continuing to increase, the moisture currently located in forest biomes will dry up.
1546: 707: 2203:. Climate change represents long-term changes in temperature and average weather patterns. This leads to a substantial increase in both the frequency and the intensity of 601: 586: 2575: 344: 3192: 4263:
Climate Change 2021: The Physical Science Basis. Working Group I contribution to the WGI Sixth Assessment Report of the Intergovernmental Panel on Climate Change
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Parmesan, C., M.D. Morecroft, Y. Trisurat, R. Adrian, G.Z. Anshari, A. Arneth, Q. Gao, P. Gonzalez, R. Harris, J. Price, N. Stevens, and G.H. Talukdarr, 2022:
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Intertidal levels: The wetness gradient of areas that are exposed to alternating water and dryness with intensities that vary by location from high to low tide
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Trends in the diversity of structure follow trends in species diversity; alpha and beta species diversities decrease from favorable to extreme environments.
4357: 687: 5577: 4289:"Predicting species distribution and abundance responses to climate change: why it is essential to include biotic interactions across trophic levels" 3109:
Olson, D. M. & E. Dinerstein (1998). The Global 200: A representation approach to conserving the Earth's most biologically valuable ecoregions.
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provide an alternative view of the terrestrial biosphere based on global patterns of sustained direct human interaction with ecosystems, including
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Whittaker based his approach on theoretical assertions and empirical sampling. He had previously compiled a review of biome classifications.
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on other climate characteristics (subarctic, warm temperate, hot temperate, and subtropical; marine and continental; lowland and mountain).
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and cracks, kilometers beneath the surface, has only recently been discovered, and does not fit well into most classification schemes.
3359: 3926:"Future projections for terrestrial biomes indicate widespread warming and moisture reduction in forests up to 2100 in South America" 3145: 2193:
system. A shift of 1 or 100% (darker colours) indicates that the region has fully moved into a completely different biome zone type.
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Anjos, Luciano J. S.; Barreiros de Souza, Everaldo; Amaral, Calil Torres; Igawa, Tassio Koiti; Mann de Toledo, Peter (2021-01-01).
3023:, 1st ed., Ulmer, Stuttgart, Germany, 1988, 488 pp.; 2nd ed., 1995, 535 pp.; 3rd ed., 2002; 4th ed., 2008; 5th ed., 2016. Transl.: 516:
patterns to relate communities to climate on a worldwide scale. Whittaker considered four main ecoclines in the terrestrial realm.
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being used when plant species are considered), or also as synonym of the "morphoclimatic and phytogeographical domain" of
4343:. Cambridge University Press, Cambridge, United Kingdom and New York, NY, USA, pp. 257-260 |doi=10.1017/9781009325844.004 3801:
Gobiet, Andreas; Kotlarski, Sven; Beniston, Martin; Heinrich, Georg; Rajczak, Jan; Stoffel, Markus (September 15, 2014).
2174: 1343:(Olson & Dinerstein, 1998, etc.). Each ecoregion is characterized by a main biome (also called major habitat type). 2283: 17: 4497: 3459: 3299: 2610: 2483: 2219:, half were found to have shifted their distribution to higher latitudes or elevations in response to climate change. 1564: 546:
Each growth-form (i.e. grasses, shrubs, etc.) has its characteristic place of maximum importance along the ecoclines.
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for 2081–2100. Top row is low emissions scenario, bottom row is high emissions scenario. Biomes are classified with
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Conditions générales de la vie dans les mers et principes de distribution des organismes marins: Année Biologique
1853: 1558: 3864:"Surface air temperature variability and trends in the Arctic: new amplification assessment and regionalisation" 3185:
Spalding, M. D. et al. (2007). Marine ecoregions of the world: a bioregionalization of coastal and shelf areas.
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Along these gradients, Whittaker noted several trends that allowed him to qualitatively establish biome-types:
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3.6% of land area will experience climates that are completely new or unusual. An example of a biome shift is
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The applicability of the realms scheme above - based on Udvardy (1975)—to most freshwater taxa is unresolved.
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General conditions of life in the seas and principles of distribution of marine organisms: Biological Year
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Whittaker's distinction between biome and formation can be simplified: formation is used when applied to
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is the collection of bacteria, viruses, and other microorganisms that are present on or in a human body.
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Biome-type: grouping of convergent biomes or formations of different continents, defined by physiognomy.
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The same growth forms may be dominant in similar environments in widely different parts of the world.
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The ecological systems of the continents (biogeosphere). Principles of their outline with examples
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Die ökologischen Systeme der Kontinente (Biogeosphäre). Prinzipien ihrer Gliederung mit Beispielen
1869:, vegetation belt between 1,100 and 1,450 m, Oromediterranean zone, nemoral zone (temperate zone) 5642: 5586: 4843: 2813: 2405: 2275: 2186: 1482: 4045:
Marcolla, Barbara; Migliavacca, Mirco; Rödenbeck, Christian; Cescatti, Alessandro (2020-04-30).
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The distribution of vegetation types as a function of mean annual temperature and precipitation.
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is a mix of organisms that coexist in a defined space on a much smaller scale. For example, the
4384: 2230: 1358: 487: 211: 156: 31: 4188: 3978:"Global tree-ring analysis reveals rapid decrease in tropical tree longevity with temperature" 3862:
Johannessen, Ola M.; Kuzmina, Svetlana I.; Bobylev, Leonid P.; Miles, Martin W. (2016-12-01).
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Box, E.O. & Fujiwara, K. (2005). Vegetation types and their broad-scale distribution. In:
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Rull, Valentí (2020). "Organisms: adaption, extinction, and biogeographical reorganizations".
5690: 5119: 4828: 4388: 4189:"Summary for Policymakers — Special Report on the Ocean and Cryosphere in a Changing Climate" 3688:
Stevens, Nicola; Lehmann, Caroline E. R.; Murphy, Brett P.; Durigan, Giselda (January 2017).
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Terrestrial biomes of the world according to Olson et al. and used by the WWF and Global 200.
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The gradient runs from favorable to the extreme, with corresponding changes in productivity.
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Kummu, Matti; Heino, Matias; Taka, Maija; Varis, Olli; Viviroli, Daniel (21 May 2021).
4096: 4022: 3977: 3957: 3903: 3778: 3741: 3665: 3620: 3596: 3536: 3465: 3416: 3325: 3089: 3000: 2967: 2873: 2769: 2740: 2690: 2533: 2374: 2365: 1580: 1403: 1391: 1112:(his concept of ecozone is more similar to the concept of biome than to the concept of 758:, provides a comprehensive coverage of the major "ecosystem types or biomes" on Earth: 640: 591: 250: 221: 4414: 4371: 4047:"Patterns and trends of the dominant environmental controls of net biome productivity" 3556:"Multivariate climate departures have outpaced univariate changes across global lands" 3491:"Past and future global transformation of terrestrial ecosystems under climate change" 5735: 5622: 5511: 5488: 5408: 5375: 4931: 4853: 4782: 4772: 4695: 4329: 4318: 4235: 4153: 4100: 4088: 4027: 4009: 3961: 3949: 3907: 3895: 3844: 3783: 3765: 3719: 3670: 3652: 3601: 3583: 3528: 3520: 3469: 3455: 3420: 3295: 3044: 2717: 2698: 2686: 2606: 2516: 2479: 2308: 2279: 1981: 1692: 1457: 1409: 1397: 943: 501: 138: 3829: 3802: 3740:
De Boeck, Hans J.; Hiltbrunner, Erika; Jentsch, Anke; Vandvik, Vigdis (2019-03-28).
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Bailey, R. G. (1989). "Explanatory supplement to ecoregions map of the continents".
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Whittaker, Robert H. (January–March 1962). "Classification of Natural Communities".
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Low, evergreen vegetation, without trees, growing over permanently frozen soils
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Walter's Vegetation of the Earth: The Ecological Systems of the Geo-Biosphere
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is already now altering biomes, adversely affecting terrestrial and marine
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ZB V. Warm temperate, occasional frost, often with summer rainfall maximum
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Abatzoglou, John T.; Dobrowski, Solomon Z.; Parks, Sean A. (2020-03-03).
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is used in a different manner. In German literature, particularly in the
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Williams, John W.; Jackson, Stephen T.; Kutzbach, John E. (2007-04-03).
102:. Biomes may span more than one continent. A biome encompasses multiple 5614: 5365: 5332: 5307: 5189: 5109: 5019: 4899: 4675: 4083: 3621:"Projected distributions of novel and disappearing climates by 2100 AD" 2869: 2773: 2537: 2378: 2181: 2124: 2022: 1888: 1844: 1816: 1810: 1783: 1777: 1750: 1744: 1721: 1632: 1626: 1605: 1347: 1340: 1326: 1091: 472: 441: 289: 284:
classified climates based on the biological effects of temperature and
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Philosophical Transactions of the Royal Society B: Biological Sciences
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Van der Putten, Wim H.; Macel, Mirka; Visser, Marcel E. (2010-07-12).
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that has formed in response to its physical environment and regional
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The Ecozones of the World: The Ecological Divisions of the Geosphere
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versus average annual temperature (y-axis) to classify biome-types.
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Chapter 2: Terrestrial and Freshwater Ecosystems and Their Services
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M310 Tropical/Subtropical Steppe Division – Mountain Provinces
961: 513: 417: 374: 285: 3923: 3690:"Savanna woody encroachment is widespread across three continents" 266: 5390: 5267: 5237: 5053: 5034: 4623: 3739: 2236: 2027: 2002: 1872: 1754: 1662: 1644: 1335:(WWF) developed a scheme that divided the world's land area into 1031: 677: 436: 293: 198: 107: 99: 83: 3269:] (in French). Vol. 2. pp. 559–587. Archived from 5257: 5124: 5104: 4597: 4487: 4358:
The Physical Environment: an Introduction to Physical Geography
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Tropical and subtropical grasslands, savannas, and shrublands
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One way of mapping terrestrial (land) biomes around the world
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Rockström, Johan; Steffen, Will; Noone, Kevin (2017-12-31),
1583:(temperate warm, semihumid to semiarid with winter rainfall) 1014:(drought-adapted), frost-sensitive shrublands and woodlands 175:
added the climatic and soil aspects to the idea, calling it
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University of California Museum of Paleontology Berkeley's
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follows. For instance, out of 4000 species analyzed by the
1614: 987: 490:: a major kind of community of plants on a given continent. 449: 67: 61: 27:
Biogeographical unit with a particular biological community
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Climate Change 2022: Impacts, Adaptation and Vulnerability
3487: 3208:"Freshwater Ecoregions of the World: Major Habitat Types" 2185:
Predicated changes for Earth's biomes under two different
1561:(temperate to tropical, fresh or brackish water inundated) 4286: 2333: 1313:
Olson & Dinerstein (1998) biomes for WWF / Global 200
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within its boundaries. It can also comprise a variety of
4952: 3553: 2245: – Concept was developed by C. Hart Merriam in 1889 1365:, nn is the biome number, NN is the individual number). 1214:
M210 Warm Continental Division – Mountain Provinces
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biome, consisting entirely of microscopic life in rock
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M220 Hot Continental Division – Mountain Provinces
2995:] (in German). Vol. 1, Grundlagen. Stuttgart. 2787:
Pomeroy, Lawrence R.; Alberts, James J., eds. (1988).
2509:"The use and abuse of vegetational terms and concepts" 1604:
According to the WWF, the following are classified as
1361:, there is a specific EcoID, format XXnnNN (XX is the 1354:
identified by the WWF as priorities for conservation.
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The principal biomes of the world by Kendeigh (1961):
30:"Biota (ecology)" redirects here. For other uses, see 5662: 3433: 3138:
ecoregions of the world: a new map of life on Earth.
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Biogeography: an ecological and evolutionary approach
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Fitossociologia no Brasil: métodos e estudos de caso.
2361:"The development and structure of biotic communities" 2165:, which can change grass savanna into shrub savanna. 1269:
M260 Mediterranean Division – Mountain Provinces
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Temperate evergreen forest, somewhat frost-sensitive
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Whittaker used what he called "gradient analysis" of
185:(1964–74) projects popularized the concept of biome. 2233: – Systems that categorize the world's climates 1780:(426), each characterized by a main biome type (12) 1747:(867), each characterized by a main biome type (14) 1339:(called "ecozones" in a BBC scheme), and these into 1068:, cold temperate with cool summers and long winters 1022:
Yellow or red forest soils, slightly podsolic soils
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Desert vegetation with considerable exposed surface
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Formation-type: a grouping of convergent formations.
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Key definitions for understanding Whittaker's scheme
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to the average conditions that predominate in them.
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M230 Subtropical Division – Mountain Provinces
70: 55: 4454: 1207:210 Warm Continental Division (Köppen: portion of 1053:, arid, with warm or hot summers and cold winters 745: 508:Whittaker's parameters for classifying biome-types 3715:20.500.11820/ff572887-5c50-4c25-8b65-a9ce5bd8ea2a 3378: 3376: 2669:"Review of plant biogeographic studies in Brazil" 2329:"Finally, A Map Of All The Microbes On Your Body" 2207:. As a region's climate changes, a change in its 1217:220 Hot Continental Division (Köppen: portion of 1197:M130 Subarctic Division – Mountain Provinces 911: 280:In 1947, the American botanist and climatologist 5722: 3735: 3733: 3027:. Berlin: Springer-Verlag, 1995; 2nd ed., 2005, 2889: 2887: 2590: 2588: 1966:(not covered yet by the Global 200/WWF scheme): 1595:(subtropical and tropical, salt water inundated) 1511:Tropical and subtropical moist broadleaf forests 1431: 3983:Proceedings of the National Academy of Sciences 3626:Proceedings of the National Academy of Sciences 2301:Quaternary Ecology, Evolution, and Biogeography 1661:Tropical and subtropical floodplain rivers and 1247:250 Prairie Division (Köppen: arid portions of 946:, always moist, little temperature seasonality 3869:Tellus A: Dynamic Meteorology and Oceanography 3436:""A Safe Operating Space for Humanity" (2009)" 3373: 3352:"What is the Endolithic Biome? (with picture)" 2786: 2468:Cox, C. B.; Moore, P.D.; Ladle, R. J. (2016). 1553:Temperate grasslands, savannas, and shrublands 1517:Tropical and subtropical dry broadleaf forests 1244:M240 Marine Division – Mountain Provinces 1187:M120 Tundra Division – Mountain Provinces 1074:Evergreen, frost-hardy, needle-leaved forest ( 5571: 4968: 4440: 4038: 3919: 3917: 3730: 2986: 2884: 2666: 2594: 2585: 2572:. Blackwell Scientific, Oxford. pp. 106–128, 2467: 2270:Bowman, William D.; Hacker, Sally D. (2021). 2251: – Living and non-living things on Earth 1227:230 Subtropical Division (Köppen: portion of 1087:, short, cool summers and long, cold winters 3133: 3131: 3129: 3105: 3103: 2269: 2151: 4588:Mediterranean forests, woodlands, and scrub 2812:. Philadelphia: Saunders Co. Archived from 1577:Mediterranean forests, woodlands, and scrub 1523:Tropical and subtropical coniferous forests 1150: 1103: 618:Subarctic-subalpine needle-leaved forests ( 305:The principal biome-types by Allee (1949): 300: 196:terminology, the term is used similarly as 64: 5578: 5564: 4975: 4961: 4447: 4433: 4396:Global Currents and Terrestrial Biomes Map 3914: 3005:: CS1 maint: location missing publisher ( 2972:: CS1 maint: location missing publisher ( 2394: 2392: 1346:This classification is used to define the 1037:Forest brown earths and grey forest soils 4312: 4147: 4082: 4072: 4021: 4003: 3943: 3889: 3838: 3828: 3777: 3759: 3713: 3664: 3646: 3595: 3514: 3410: 3283: 3126: 3100: 2893: 2845: 2736: 2651: 2418: 1005:, winter rainy season and summer drought 350: 4360:. University of Wisconsin-Stevens Point. 3179: 2826: 2438:Vol. 1. Viçosa: Editora UFV. pp. 44–85. 2398: 2358: 2180: 1727: 1316: 1285:320 Tropical/Subtropical Desert Division 1279:310 Tropical/Subtropical Steppe Division 1034:, moderate climate with winter freezing 798:Temperate Broad-Leaved Evergreen Forests 453: 450:Whittaker (1962, 1970, 1975) biome-types 265: 36: 3013: 2711: 2633: 2506: 2389: 2058:Humans have altered global patterns of 2047: 1504: 113:While a biome can cover small areas, a 14: 5723: 3390:Communications Earth & Environment 3321:"The Next Frontier: The Great Indoors" 3318: 3258: 3061: 2953: 2636:"The Brazilian cerrado is not a biome" 1599: 1090:Tundra humus soils with solifluction ( 205:In the Brazilian literature, the term 5585: 5559: 4956: 4428: 4276:from the original on 4 November 2021. 3155: 2805: 1858:temperate broadleaf and mixed forests 1529:Temperate broadleaf and mixed forests 1525:(tropical and subtropical, semihumid) 1519:(tropical and subtropical, semihumid) 1331:A team of biologists convened by the 877:Ecosystems of the Continental Shelves 5507:Geology of solar terrestrial planets 4572:Grasslands, savannas, and shrublands 4534:Grasslands, savannas, and shrublands 3211:"Freshwater Ecoregions of the World" 2674:Journal of Systematics and Evolution 2298: 2135: 1549:(tropical and subtropical, semiarid) 1262:260 Mediterranean Division (Köppen: 292:under the assumption that these two 188:However, in some contexts, the term 2928: 2175:Effects of climate change on biomes 2118: 1800:Biogeographic realms (marine) (12) 822:Managed Terrestrial Ecosystems 767:Natural Terrestrial Ecosystems 24: 4350: 3333:from the original on June 14, 2018 2987:Walter, H.; Breckle, S-W. (1991). 2667:Fiaschi, P.; Pirani, J.R. (2009). 2595:Walter, H.; Breckle, S-W. (2002). 1922:Pruvot (1896) zones or "systems": 1686: 1108:Schultz (1988, 2005) defined nine 868:Intertidal and Littoral Ecosystems 783:Temperate Deserts and Semi-Deserts 728:Temperate fresh-water swamp forest 232: 209:is sometimes used as a synonym of 137:The term was suggested in 1916 by 25: 5747: 4498:Montane grasslands and shrublands 4364: 4233: 1565:Montane grasslands and shrublands 1513:(tropical and subtropical, humid) 1059:Grasslands and temperate deserts 795:Heathlands and Related Shrublands 723:Tropical fresh-water swamp forest 262:Holdridge (1947, 1964) life zones 5708: 5696: 5684: 5672: 5540: 5539: 5043: 3808:Science of the Total Environment 3236:"Marine Ecoregions of the World" 2687:10.1111/j.1759-6831.2009.00046.x 2173:This section is an excerpt from 1911: 1877:Oreoherzogio-Abietetum illyricae 1643:Temperate floodplain rivers and 1190:130 Subarctic Division (Köppen: 780:Mires: Swamp, Bog, Fen, and Moor 529:Temperature gradient by latitude 526:Temperature gradient by altitude 183:International Biological Program 51: 5248:Human impact on the environment 4614:Flooded grasslands and savannas 4280: 4227: 4201: 4181: 4164: 3968: 3931:Global Ecology and Conservation 3855: 3830:10.1016/j.scitotenv.2013.07.050 3794: 3681: 3612: 3547: 3481: 3427: 3362:from the original on 2017-03-07 3344: 3319:Zimmer, Carl (March 19, 2015). 3312: 3287:Ecological Geography of the Sea 3277: 3252: 3228: 3202: 3055: 3033: 2980: 2947: 2922: 2906: 2839: 2820: 2799: 2780: 2730: 2705: 2660: 2653:10.1590/S1676-06032011000100001 2627: 2420:10.1590/S0102-33062006000100002 2341:from the original on 2018-04-16 1906: 1854:Dinaric Mountains mixed forests 1559:Flooded grasslands and savannas 1537:(temperate, humid to semihumid) 1136:subtropics with year-round rain 886:Managed Aquatic Ecosystems 810:Tropical Rain Forest Ecosystems 804:Hot Deserts and Arid Shrublands 746:Goodall (1974–) ecosystem types 241:A 1978 study on North American 5077:Climate variability and change 4927:Ecological land classification 4456:Biogeographic regionalisations 2558: 2500: 2461: 2427: 2352: 2321: 2292: 2263: 912:Walter (1976, 2002) zonobiomes 851:Inland Aquatic Ecosystems 816:Ecosystems of Disturbed Ground 557: 141:, originally as a synonym for 132: 94:. It consists of a biological 13: 1: 5502:Evolution of the Solar System 2962:] (in German). Stuttgart. 2894:Whittaker, Robert H. (1975). 2789:Concepts of Ecosystem Ecology 2403:[The biome concept]. 2256: 2032:Coastal lagoons/Atoll lagoons 1589:(temperate to tropical, arid) 1291:340 Temperate Desert Division 1288:330 Temperate Steppe Division 1237:240 Marine Division (Köppen: 801:Mediterranean-Type Shrublands 5253:Evolutionary history of life 4593:Deserts and xeric shrublands 4260:"Summary for Policymakers". 4140:10.1016/j.oneear.2021.04.017 2809:Principles of animal ecology 2217:IPCC Sixth Assessment Report 2131:Habitat § Microhabitats 2094:Major anthropogenic biomes: 1587:Deserts and xeric shrublands 1535:Temperate coniferous forests 1372:(terrestrial and freshwater) 1145:tropics with year-round rain 880:Ecosystems of the Deep Ocean 570:Tropical seasonal rainforest 7: 4834:Temperate Northern Atlantic 4524:Broadleaf and mixed forests 4403:(archived 22 February 2011) 4356:Ritter, Michael E. (2005). 3945:10.1016/j.gecco.2020.e01441 2935:. Vol. 36. Amsterdam: 2223: 1567:(alpine or montane climate) 1448:Temperate Northern Atlantic 1133:subtropics with winter rain 1008:Mediterranean brown earths 874:Estuaries and Enclosed Seas 855:River and Stream Ecosystems 789:Temperate Deciduous Forests 708:Cool temperate desert scrub 10: 5752: 5041: 4937:Vegetation classifications 4839:Temperate Northern Pacific 4420:World Map of Desert Biomes 3747:Frontiers in Plant Science 3580:10.1038/s41598-020-60270-5 3452:10.12987/9780300188479-042 3412:10.1038/s43247-021-00270-z 3065:Environmental Conservation 2914:Communities and Ecosystems 2896:Communities and Ecosystems 2750:British Ecological Society 2716:. Springer. pp. 2–3. 2619:on 2016-11-27 – via 2492:on 2016-11-26 – via 2172: 2128: 2122: 2051: 1915: 1691:Biomes of the coastal and 1453:Temperate Northern Pacific 1324: 1203:200 Humid Temperate Domain 1139:dry tropics and subtropics 890:Managed Aquatic Ecosystems 750:The multi-authored series 602:Temperate evergreen forest 597:Temperate deciduous forest 587:Temperate giant rainforest 523:Climatic moisture gradient 365:Temperate deciduous forest 273: 29: 5593: 5525: 5487: 5444: 5389: 5331: 5266: 5213: 5143: 5095: 5052: 4990: 4982: 4914: 4872: 4849:Temperate Southern Africa 4801: 4745: 4734: 4714: 4641: 4606: 4580: 4542: 4511: 4480: 4471: 4462: 4413:NASA's Earth Observatory 3891:10.3402/tellusa.v68.28234 3086:10.1017/S0376892900009711 2912:Whittaker, R. H. (1970). 2714:The ecozones of the world 2474:(9th ed.). Hoboken: 2448:. Earlier version, 2003, 2152:Effects of climate change 1706:Temperate shelves and sea 1667:Tropical and subtropical 1655:Tropical and subtropical 1488:Temperate Southern Africa 1297:400 Humid Tropical Domain 1162:classification system of 899:Underground Ecosystems 763:Terrestrial Ecosystems 5512:Location in the Universe 5454:Antarctic/Southern Ocean 5153:List of sovereign states 4859:Tropical Eastern Pacific 4809:Antarctic/Southern Ocean 2831:. Englewood Cliffs, NJ: 2712:Schultz, Jürgen (1995). 2399:Coutinho, L. M. (2006). 2359:Clements, F. E. (1917). 2187:climate change scenarios 2163:woody plant encroachment 1478:Tropical Eastern Pacific 1151:Bailey (1989) ecoregions 1142:tropics with summer rain 1104:Schultz (1988) eco-zones 971:Tropical seasonal forest 967:Red clays or red earths 901: 888: 866: 853: 838:Bioindustrial Ecosystems 824: 345:Minor terrestrial biomes 301:Allee (1949) biome-types 4880:Biogeographic provinces 4844:Temperate South America 4562:Moist broadleaf forests 4380:(archived 11 July 2011) 4074:10.5194/bg-17-2365-2020 4005:10.1073/pnas.2003873117 3761:10.3389/fpls.2019.00347 3648:10.1073/pnas.0606292104 3516:10.1126/science.aan5360 2932:Ecosystems of the World 2827:Kendeigh, S.C. (1961). 2406:Acta Botanica Brasilica 2276:Oxford University Press 1999:(trades and westerlies) 1943:Longhurst (1998) biomes 1483:Temperate South America 1304:420 Rainforest Division 1056:Chernozems to serozems 949:Equatorial brown clays 777:Polar and Alpine Tundra 752:Ecosystems of the World 163:(used in opposition to 4410:(archived 6 July 2017) 4305:10.1098/rstb.2010.0037 3284:Longhurst, A. (1998). 2929:Goodall, D. W. (ed.). 2916:. Toronto, pp. 51–64, 2634:Batalha, M.A. (2011). 2507:Tansley, A.G. (1935). 2231:Climate classification 2205:extreme weather events 2194: 1856:(PA0418); biome type: 1673:Xeric freshwaters and 1432:Biogeographic realms ( 1359:terrestrial ecoregions 1322: 993:Serosemes, sierozemes 937:Zonal vegetation type 916:The eponymously named 864:Marine Ecosystems 774:Dry Coastal Ecosystems 771:Wet Coastal Ecosystems 459: 351:Kendeigh (1961) biomes 271: 212:biogeographic province 42: 5233:Biogeochemical cycles 5158:dependent territories 4942:Zoogeographic regions 4900:Global 200 ecoregions 4829:Temperate Australasia 4567:Dry broadleaf forests 4389:Encyclopedia of Earth 3694:Global Change Biology 3444:Yale University Press 3021:Die Ökozonen der Erde 2476:John Wiley & Sons 2401:"O conceito de bioma" 2184: 2129:Further information: 2052:Further information: 1916:Further information: 1728:Summary of the scheme 1555:(temperate, semiarid) 1493:Temperate Australasia 1320: 847:Aquatic Ecosystems 835:Greenhouse Ecosystems 829:Field Crop Ecosystems 703:Warm-temperate desert 457: 269: 40: 5643:Biomolecular complex 5281:Computer cartography 5005:Prebiotic atmosphere 4922:Altitudinal zonation 4864:Western Indo-Pacific 4824:Eastern Indo-Pacific 4819:Central Indo-Pacific 3446:, pp. 491–505, 3440:The Future of Nature 2993:Ecology of the Earth 2900:MacMillan Publishing 2806:Allee, W.C. (1949). 2278:. pp. H3–1–51. 2191:Holdridge life zones 2069:Anthropogenic biomes 2048:Anthropogenic biomes 1772:Biogeographic realms 1739:Biogeographic realms 1541:Boreal forests/taiga 1505:Biomes (terrestrial) 1473:Eastern Indo-Pacific 1468:Central Indo-Pacific 1463:Western Indo-Pacific 1370:Biogeographic realms 1337:biogeographic realms 1301:410 Savanna Division 977:, scrub, or savanna 858:Lakes and Reservoirs 832:Tree Crop Ecosystems 713:Arctic-alpine desert 666:subalpine-needleleaf 646:Temperate shrublands 276:Holdridge life zones 247:logistic correlation 5424:Geologic time scale 5145:Culture and society 5000:Atmosphere of Earth 4895:Lists of ecoregions 4552:Tropical rainforest 4408:Major Habitat Types 4299:(1549): 2025–2034. 4213:National Geographic 4132:2021OEart...4..720K 4065:2020BGeo...17.2365M 3996:2020PNAS..11733358M 3990:(52): 33358–33364. 3882:2016TellA..6828234J 3821:2014ScTEn.493.1138G 3639:2007PNAS..104.5738W 3572:2020NatSR..10.3891A 3507:2018Sci...361..920N 3403:2021ComEE...2..198D 3259:Pruvot, G. (1896). 3078:1989EnvCo..16..307B 2954:Walter, H. (1976). 2862:1962BotRv..28....1W 2758:1978JEcol..66..573S 2249:Natural environment 2054:Anthropogenic biome 1886:Plant: Silver fir ( 1841:Biogeographic realm 1600:Biomes (freshwater) 1581:sclerophyll forests 1363:biogeographic realm 1333:World Wildlife Fund 1158:nearly developed a 1127:humid mid-latitudes 1121:polar/subpolar zone 1044:, temperate forest 986:, highly seasonal, 954:tropical rainforest 683:Temperate grassland 671:subalpine-broadleaf 565:Tropical rainforest 426:Balanoid-gastropod- 217:species composition 215:, an area based on 169:species composition 80:geographical region 5419:Geological history 5293:Geodetic astronomy 4932:Floristic kingdoms 4696:Hydrothermal vents 4557:Coniferous forests 4519:Coniferous forests 4372:The World's Biomes 4234:Witze, Alexandra. 3840:20.500.11850/87298 3560:Scientific Reports 3326:The New York Times 3195:2016-10-06 at the 3172:2016-10-06 at the 3148:2012-09-17 at the 3119:2016-10-07 at the 3111:Conservation Biol. 2870:10.1007/BF02860872 2741:Journal of Ecology 2578:2016-08-28 at the 2570:Vegetation ecology 2566:van der Maarel, E. 2454:2016-08-27 at the 2444:2016-09-24 at the 2366:Journal of Ecology 2195: 1982:Hydrothermal vents 1741:(terrestrial) (8) 1543:(subarctic, humid) 1531:(temperate, humid) 1323: 1180:Division (Köppen: 826:Managed Grasslands 792:Natural Grasslands 786:Coniferous Forests 641:Temperate woodland 592:Montane rainforest 460: 272: 251:evapotranspiration 222:floristic province 43: 18:Major habitat type 5660: 5659: 5587:Hierarchy of life 5553: 5552: 5489:Planetary science 5409:Extremes on Earth 5376:Signal processing 4950: 4949: 4910: 4909: 4854:Tropical Atlantic 4730: 4729: 4637: 4636: 4529:Deciduous forests 3706:10.1111/gcb.13409 3633:(14): 5738–5742. 3501:(6405): 920–923. 3045:US Forest Service 2989:Ökologie der Erde 2723:978-3-540-28527-4 2409:(in Portuguese). 2314:978-0-12-820473-3 2136:Endolithic biomes 2098:Dense settlements 2077:human settlements 1774:(freshwater) (8) 1693:continental shelf 1458:Tropical Atlantic 1130:dry mid-latitudes 1101: 1100: 1040:Frost-resistant, 807:Tropical Savannas 688:Alpine grasslands 502:plant communities 370:Coniferous forest 255:solar irradiation 245:found a positive 16:(Redirected from 5743: 5713: 5712: 5701: 5700: 5699: 5689: 5688: 5687: 5677: 5676: 5668: 5653: 5580: 5573: 5566: 5557: 5556: 5543: 5542: 5436:History of Earth 5087:Paleoclimatology 5047: 4977: 4970: 4963: 4954: 4953: 4743: 4742: 4478: 4477: 4469: 4468: 4449: 4442: 4435: 4426: 4425: 4392: 4344: 4333: 4327: 4326: 4316: 4284: 4278: 4277: 4275: 4268: 4257: 4251: 4250: 4248: 4246: 4231: 4225: 4224: 4222: 4220: 4209:"Climate Change" 4205: 4199: 4198: 4196: 4195: 4185: 4179: 4178: 4176: 4168: 4162: 4161: 4151: 4111: 4105: 4104: 4086: 4076: 4059:(8): 2365–2379. 4042: 4036: 4035: 4025: 4007: 3972: 3966: 3965: 3947: 3921: 3912: 3911: 3893: 3859: 3853: 3852: 3842: 3832: 3798: 3792: 3791: 3781: 3763: 3737: 3728: 3727: 3717: 3685: 3679: 3678: 3668: 3650: 3616: 3610: 3609: 3599: 3551: 3545: 3544: 3518: 3485: 3479: 3478: 3477: 3476: 3431: 3425: 3424: 3414: 3380: 3371: 3370: 3368: 3367: 3348: 3342: 3341: 3339: 3338: 3316: 3310: 3309: 3281: 3275: 3274: 3256: 3250: 3249: 3244:. Archived from 3232: 3226: 3225: 3223: 3222: 3213:. Archived from 3206: 3200: 3183: 3177: 3159: 3153: 3142:51(11):933–938, 3135: 3124: 3107: 3098: 3097: 3059: 3053: 3052: 3047:. Archived from 3037: 3031: 3017: 3011: 3010: 3004: 2996: 2984: 2978: 2977: 2971: 2963: 2951: 2945: 2944: 2939:. Archived from 2926: 2920: 2910: 2904: 2903: 2891: 2882: 2881: 2849:Botanical Review 2843: 2837: 2836: 2824: 2818: 2817: 2803: 2797: 2796: 2784: 2778: 2777: 2734: 2728: 2727: 2709: 2703: 2702: 2697:. Archived from 2664: 2658: 2657: 2655: 2640:Biota Neotropica 2631: 2625: 2624: 2615:. Archived from 2592: 2583: 2562: 2556: 2555: 2553: 2552: 2546: 2540:. Archived from 2513: 2504: 2498: 2497: 2488:. Archived from 2465: 2459: 2431: 2425: 2424: 2422: 2396: 2387: 2386: 2381:. Archived from 2356: 2350: 2349: 2347: 2346: 2325: 2319: 2318: 2296: 2290: 2289: 2274:(5th ed.). 2267: 2119:Microbial biomes 1805:Marine provinces 1675:endorheic basins 1172:100 Polar Domain 1156:Robert G. Bailey 934:Zonal soil type 928: 927: 923:vegetation types 756:David W. Goodall 400:Tropical savanna 320:Deciduous forest 282:Leslie Holdridge 153:phytophysiognomy 144:biotic community 119:human microbiome 78:) is a distinct 77: 76: 73: 72: 69: 66: 63: 60: 57: 21: 5751: 5750: 5746: 5745: 5744: 5742: 5741: 5740: 5721: 5720: 5719: 5707: 5697: 5695: 5685: 5683: 5671: 5663: 5661: 5656: 5597: 5589: 5584: 5554: 5549: 5521: 5483: 5440: 5431:Geologic record 5385: 5371:Plate tectonics 5361:Mineral physics 5341:Earth structure 5327: 5262: 5209: 5139: 5091: 5048: 5039: 4986: 4981: 4951: 4946: 4906: 4868: 4797: 4737: 4726: 4722:Endolithic zone 4710: 4643: 4633: 4602: 4576: 4544: 4538: 4507: 4473: 4458: 4453: 4415:Mission: Biomes 4401:WorldBiomes.com 4383: 4367: 4353: 4351:Further reading 4348: 4347: 4334: 4330: 4285: 4281: 4273: 4266: 4259: 4258: 4254: 4244: 4242: 4232: 4228: 4218: 4216: 4215:. 28 March 2019 4207: 4206: 4202: 4193: 4191: 4187: 4186: 4182: 4174: 4170: 4169: 4165: 4112: 4108: 4043: 4039: 3973: 3969: 3922: 3915: 3860: 3856: 3799: 3795: 3738: 3731: 3686: 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2133: 2127: 2121: 2056: 2050: 2045: 2018:Intertidal zone 1920: 1918:Marine habitats 1914: 1909: 1730: 1689: 1687:Biomes (marine) 1684: 1680:Oceanic islands 1602: 1507: 1502: 1438: 1426: 1374: 1329: 1315: 1160:biogeographical 1153: 1148: 1106: 918:Heinrich Walter 914: 909: 906: 903:Cave Ecosystems 896: 893: 883: 861: 844: 841: 819: 813:Wetland Forests 748: 743: 698:Tropical desert 560: 510: 475: 452: 405:Tropical forest 353: 340:Tropical forest 303: 278: 264: 235: 233:Classifications 135: 54: 50: 35: 28: 23: 22: 15: 12: 11: 5: 5749: 5739: 5738: 5733: 5718: 5717: 5705: 5693: 5691:Earth sciences 5681: 5658: 5657: 5655: 5654: 5594: 5591: 5590: 5583: 5582: 5575: 5568: 5560: 5551: 5550: 5548: 5547: 5537: 5532: 5526: 5523: 5522: 5520: 5519: 5514: 5509: 5504: 5499: 5493: 5491: 5485: 5484: 5482: 5481: 5476: 5471: 5466: 5464:Atlantic Ocean 5461: 5456: 5450: 5448: 5442: 5441: 5439: 5438: 5433: 5428: 5427: 5426: 5416: 5411: 5406: 5401: 5395: 5393: 5387: 5386: 5384: 5383: 5378: 5373: 5368: 5363: 5358: 5353: 5348: 5346:Fluid dynamics 5343: 5337: 5335: 5329: 5328: 5326: 5325: 5320: 5318:Geopositioning 5315: 5313:Remote Sensing 5310: 5305: 5300: 5295: 5290: 5285: 5284: 5283: 5272: 5270: 5264: 5263: 5261: 5260: 5255: 5250: 5245: 5240: 5235: 5230: 5225: 5219: 5217: 5211: 5210: 5208: 5207: 5202: 5197: 5192: 5187: 5182: 5177: 5172: 5167: 5162: 5161: 5160: 5149: 5147: 5141: 5140: 5138: 5137: 5132: 5127: 5122: 5117: 5112: 5107: 5101: 5099: 5093: 5092: 5090: 5089: 5084: 5079: 5074: 5072:Climate change 5069: 5067:Energy balance 5064: 5062:Climate system 5058: 5056: 5050: 5049: 5042: 5040: 5038: 5037: 5032: 5027: 5022: 5017: 5012: 5007: 5002: 4996: 4994: 4988: 4987: 4980: 4979: 4972: 4965: 4957: 4948: 4947: 4945: 4944: 4939: 4934: 4929: 4924: 4918: 4916: 4912: 4911: 4908: 4907: 4905: 4904: 4903: 4902: 4897: 4887: 4882: 4876: 4874: 4870: 4869: 4867: 4866: 4861: 4856: 4851: 4846: 4841: 4836: 4831: 4826: 4821: 4816: 4811: 4805: 4803: 4799: 4798: 4796: 4795: 4790: 4785: 4780: 4775: 4770: 4765: 4760: 4755: 4749: 4747: 4740: 4736:Biogeographic 4732: 4731: 4728: 4727: 4725: 4724: 4718: 4716: 4712: 4711: 4709: 4708: 4703: 4698: 4693: 4688: 4683: 4678: 4673: 4668: 4663: 4658: 4653: 4647: 4645: 4639: 4638: 4635: 4634: 4632: 4631: 4626: 4621: 4616: 4610: 4608: 4604: 4603: 4601: 4600: 4595: 4590: 4584: 4582: 4578: 4577: 4575: 4574: 4569: 4564: 4559: 4554: 4548: 4546: 4540: 4539: 4537: 4536: 4531: 4526: 4521: 4515: 4513: 4509: 4508: 4506: 4505: 4500: 4495: 4490: 4484: 4482: 4475: 4466: 4460: 4459: 4452: 4451: 4444: 4437: 4429: 4423: 4422: 4417: 4411: 4404: 4398: 4393: 4381: 4378:Biome Overview 4374: 4366: 4365:External links 4363: 4362: 4361: 4352: 4349: 4346: 4345: 4328: 4279: 4252: 4226: 4200: 4180: 4163: 4126:(5): 720–729. 4106: 4052:Biogeosciences 4037: 3967: 3913: 3854: 3793: 3729: 3700:(1): 235–244. 3680: 3611: 3546: 3480: 3460: 3426: 3372: 3343: 3311: 3300: 3292:Academic Press 3276: 3273:on 2016-10-18. 3251: 3248:on 2009-02-07. 3227: 3201: 3178: 3154: 3125: 3099: 3072:(4): 307–309. 3054: 3051:on 2009-01-01. 3032: 3012: 2979: 2946: 2943:on 2016-09-18. 2921: 2905: 2883: 2838: 2829:Animal ecology 2819: 2816:on 2017-10-01. 2798: 2779: 2729: 2722: 2704: 2701:on 2017-08-31. 2681:(5): 477–496. 2659: 2626: 2611: 2605:. p. 86. 2584: 2557: 2524:(3): 284–307. 2499: 2484: 2478:. p. 20. 2460: 2426: 2388: 2385:on 2016-10-07. 2351: 2320: 2313: 2307:. p. 67. 2305:Academic Press 2291: 2285:978-1605359212 2284: 2261: 2260: 2258: 2255: 2253: 2252: 2246: 2240: 2234: 2227: 2225: 2222: 2197:Climate change 2179: 2171: 2158:climate change 2156:Anthropogenic 2153: 2150: 2137: 2134: 2123:Main article: 2120: 2117: 2116: 2115: 2112: 2109: 2104: 2099: 2049: 2046: 2044: 2043: 2038: 2033: 2030: 2025: 2020: 2011: 2010:(ocean trench) 2005: 2000: 1994: 1989: 1984: 1979: 1974: 1968: 1960: 1959: 1956: 1953: 1950: 1940: 1939: 1934: 1929: 1913: 1910: 1908: 1905: 1904: 1903: 1902: 1901: 1900: 1899: 1898: 1897: 1896: 1895: 1894: 1893: 1829: 1828: 1827: 1826: 1825: 1824: 1823: 1822: 1821: 1820: 1793: 1792: 1791: 1790: 1789: 1788: 1787: 1764: 1763: 1762: 1761: 1760: 1759: 1758: 1729: 1726: 1725: 1724: 1722:Tropical coral 1719: 1713: 1707: 1704: 1688: 1685: 1683: 1682: 1677: 1671: 1665: 1659: 1657:coastal rivers 1653: 1647: 1641: 1639:coastal rivers 1635: 1629: 1623: 1617: 1610: 1601: 1598: 1597: 1596: 1590: 1584: 1574: 1568: 1562: 1556: 1550: 1544: 1538: 1532: 1526: 1520: 1514: 1506: 1503: 1501: 1500: 1498:Southern Ocean 1495: 1490: 1485: 1480: 1475: 1470: 1465: 1460: 1455: 1450: 1445: 1439: 1437: 1430: 1425: 1424: 1418: 1412: 1406: 1400: 1394: 1388: 1382: 1375: 1373: 1367: 1325:Main article: 1314: 1311: 1308: 1307: 1306: 1305: 1302: 1294: 1293: 1292: 1289: 1286: 1283: 1280: 1275:300 Dry Domain 1272: 1271: 1270: 1267: 1260: 1245: 1242: 1235: 1232: 1225: 1222: 1215: 1212: 1200: 1199: 1198: 1195: 1188: 1185: 1152: 1149: 1147: 1146: 1143: 1140: 1137: 1134: 1131: 1128: 1125: 1122: 1118: 1105: 1102: 1099: 1098: 1095: 1088: 1080: 1079: 1072: 1069: 1061: 1060: 1057: 1054: 1046: 1045: 1038: 1035: 1027: 1026: 1023: 1020: 1016: 1015: 1012:Sclerophyllous 1009: 1006: 998: 997: 994: 991: 979: 978: 968: 965: 957: 956: 950: 947: 939: 938: 935: 932: 913: 910: 908: 907: 905: 904: 900: 897: 895: 894: 892: 891: 887: 884: 882: 881: 878: 875: 872: 869: 865: 862: 860: 859: 856: 852: 848: 845: 843: 842: 840: 839: 836: 833: 830: 827: 823: 820: 818: 817: 814: 811: 808: 805: 802: 799: 796: 793: 790: 787: 784: 781: 778: 775: 772: 768: 764: 760: 747: 744: 742: 741: 738: 735: 733:Mangrove swamp 730: 725: 720: 715: 710: 705: 700: 695: 690: 685: 680: 675: 674: 673: 668: 663: 658: 653: 643: 638: 633: 628: 626:Elfin woodland 623: 616: 615: 614: 609: 599: 594: 589: 584: 583: 582: 577: 567: 561: 559: 556: 551: 550: 547: 544: 541: 538: 531: 530: 527: 524: 521: 509: 506: 498: 497: 494: 491: 485: 482: 474: 471: 451: 448: 447: 446: 445: 444: 439: 430: 424: 409: 408: 407: 402: 397: 392: 387: 382: 377: 372: 367: 352: 349: 348: 347: 342: 337: 332: 327: 322: 317: 312: 302: 299: 274:Main article: 263: 260: 234: 231: 134: 131: 82:with specific 26: 9: 6: 4: 3: 2: 5748: 5737: 5734: 5732: 5729: 5728: 5726: 5716: 5711: 5706: 5704: 5694: 5692: 5682: 5680: 5675: 5670: 5669: 5666: 5652: 5648: 5647:Macromolecule 5644: 5640: 5636: 5632: 5628: 5624: 5620: 5616: 5612: 5608: 5604: 5600: 5596: 5595: 5592: 5588: 5581: 5576: 5574: 5569: 5567: 5562: 5561: 5558: 5546: 5538: 5536: 5533: 5531: 5528: 5527: 5524: 5518: 5515: 5513: 5510: 5508: 5505: 5503: 5500: 5498: 5495: 5494: 5492: 5490: 5486: 5480: 5477: 5475: 5474:Pacific Ocean 5472: 5470: 5467: 5465: 5462: 5460: 5457: 5455: 5452: 5451: 5449: 5447: 5443: 5437: 5434: 5432: 5429: 5425: 5422: 5421: 5420: 5417: 5415: 5412: 5410: 5407: 5405: 5404:Earth science 5402: 5400: 5397: 5396: 5394: 5392: 5388: 5382: 5379: 5377: 5374: 5372: 5369: 5367: 5364: 5362: 5359: 5357: 5356:Magnetosphere 5354: 5352: 5349: 5347: 5344: 5342: 5339: 5338: 5336: 5334: 5330: 5324: 5323:Virtual globe 5321: 5319: 5316: 5314: 5311: 5309: 5306: 5304: 5301: 5299: 5296: 5294: 5291: 5289: 5288:Earth's orbit 5286: 5282: 5279: 5278: 5277: 5274: 5273: 5271: 5269: 5265: 5259: 5256: 5254: 5251: 5249: 5246: 5244: 5241: 5239: 5236: 5234: 5231: 5229: 5226: 5224: 5221: 5220: 5218: 5216: 5212: 5206: 5203: 5201: 5198: 5196: 5195:World history 5193: 5191: 5188: 5186: 5185:World economy 5183: 5181: 5178: 5176: 5173: 5171: 5168: 5166: 5163: 5159: 5156: 5155: 5154: 5151: 5150: 5148: 5146: 5142: 5136: 5135:South America 5133: 5131: 5130:North America 5128: 5126: 5123: 5121: 5118: 5116: 5113: 5111: 5108: 5106: 5103: 5102: 5100: 5098: 5094: 5088: 5085: 5083: 5080: 5078: 5075: 5073: 5070: 5068: 5065: 5063: 5060: 5059: 5057: 5055: 5051: 5046: 5036: 5033: 5031: 5028: 5026: 5023: 5021: 5018: 5016: 5013: 5011: 5008: 5006: 5003: 5001: 4998: 4997: 4995: 4993: 4989: 4985: 4978: 4973: 4971: 4966: 4964: 4959: 4958: 4955: 4943: 4940: 4938: 4935: 4933: 4930: 4928: 4925: 4923: 4920: 4919: 4917: 4913: 4901: 4898: 4896: 4893: 4892: 4891: 4888: 4886: 4883: 4881: 4878: 4877: 4875: 4871: 4865: 4862: 4860: 4857: 4855: 4852: 4850: 4847: 4845: 4842: 4840: 4837: 4835: 4832: 4830: 4827: 4825: 4822: 4820: 4817: 4815: 4812: 4810: 4807: 4806: 4804: 4800: 4794: 4791: 4789: 4786: 4784: 4781: 4779: 4776: 4774: 4771: 4769: 4766: 4764: 4761: 4759: 4756: 4754: 4751: 4750: 4748: 4744: 4741: 4739: 4733: 4723: 4720: 4719: 4717: 4713: 4707: 4706:Demersal zone 4704: 4702: 4699: 4697: 4694: 4692: 4689: 4687: 4684: 4682: 4679: 4677: 4674: 4672: 4669: 4667: 4664: 4662: 4659: 4657: 4654: 4652: 4649: 4648: 4646: 4640: 4630: 4627: 4625: 4622: 4620: 4617: 4615: 4612: 4611: 4609: 4605: 4599: 4596: 4594: 4591: 4589: 4586: 4585: 4583: 4579: 4573: 4570: 4568: 4565: 4563: 4560: 4558: 4555: 4553: 4550: 4549: 4547: 4541: 4535: 4532: 4530: 4527: 4525: 4522: 4520: 4517: 4516: 4514: 4510: 4504: 4503:Alpine tundra 4501: 4499: 4496: 4494: 4491: 4489: 4486: 4485: 4483: 4481:Polar/montane 4479: 4476: 4470: 4467: 4465: 4461: 4457: 4450: 4445: 4443: 4438: 4436: 4431: 4430: 4427: 4421: 4418: 4416: 4412: 4409: 4405: 4402: 4399: 4397: 4394: 4390: 4386: 4382: 4379: 4376:Gale/Cengage 4375: 4373: 4369: 4368: 4359: 4355: 4354: 4342: 4338: 4332: 4324: 4320: 4315: 4310: 4306: 4302: 4298: 4294: 4290: 4283: 4272: 4265: 4264: 4256: 4241: 4237: 4230: 4214: 4210: 4204: 4190: 4184: 4173: 4167: 4159: 4155: 4150: 4145: 4141: 4137: 4133: 4129: 4125: 4121: 4117: 4110: 4102: 4098: 4094: 4090: 4085: 4080: 4075: 4070: 4066: 4062: 4058: 4054: 4053: 4048: 4041: 4033: 4029: 4024: 4019: 4015: 4011: 4006: 4001: 3997: 3993: 3989: 3985: 3984: 3979: 3971: 3963: 3959: 3955: 3951: 3946: 3941: 3937: 3933: 3932: 3927: 3920: 3918: 3909: 3905: 3901: 3897: 3892: 3887: 3883: 3879: 3875: 3871: 3870: 3865: 3858: 3850: 3846: 3841: 3836: 3831: 3826: 3822: 3818: 3815:: 1138–1151. 3814: 3810: 3809: 3804: 3797: 3789: 3785: 3780: 3775: 3771: 3767: 3762: 3757: 3753: 3749: 3748: 3743: 3736: 3734: 3725: 3721: 3716: 3711: 3707: 3703: 3699: 3695: 3691: 3684: 3676: 3672: 3667: 3662: 3658: 3654: 3649: 3644: 3640: 3636: 3632: 3628: 3627: 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San Diego: 3289: 3288: 3280: 3272: 3268: 3264: 3263: 3255: 3247: 3243: 3242: 3237: 3231: 3217:on 2008-10-07 3216: 3212: 3205: 3198: 3194: 3191: 3189:57: 573–583, 3188: 3182: 3175: 3171: 3168: 3165: 3158: 3151: 3147: 3144: 3141: 3134: 3132: 3130: 3122: 3118: 3115: 3112: 3106: 3104: 3095: 3091: 3087: 3083: 3079: 3075: 3071: 3067: 3066: 3058: 3050: 3046: 3042: 3036: 3029: 3026: 3022: 3016: 3008: 3002: 2994: 2990: 2983: 2975: 2969: 2961: 2957: 2950: 2942: 2938: 2934: 2933: 2925: 2918: 2915: 2909: 2901: 2897: 2890: 2888: 2879: 2875: 2871: 2867: 2863: 2859: 2855: 2851: 2850: 2842: 2834: 2833:Prentice-Hall 2830: 2823: 2815: 2811: 2810: 2802: 2794: 2790: 2783: 2775: 2771: 2767: 2763: 2759: 2755: 2751: 2747: 2743: 2742: 2733: 2725: 2719: 2715: 2708: 2700: 2696: 2692: 2688: 2684: 2680: 2676: 2675: 2670: 2663: 2654: 2649: 2645: 2641: 2637: 2630: 2622: 2618: 2614: 2612:9783540433156 2608: 2604: 2600: 2599: 2591: 2589: 2581: 2577: 2574: 2571: 2567: 2561: 2547:on 2016-10-06 2543: 2539: 2535: 2531: 2527: 2523: 2519: 2518: 2510: 2503: 2495: 2491: 2487: 2485:9781118968581 2481: 2477: 2473: 2472: 2464: 2457: 2453: 2450: 2447: 2443: 2440: 2437: 2430: 2421: 2416: 2412: 2408: 2407: 2402: 2395: 2393: 2384: 2380: 2376: 2372: 2368: 2367: 2362: 2355: 2340: 2336: 2335: 2330: 2324: 2316: 2310: 2306: 2302: 2295: 2287: 2281: 2277: 2273: 2266: 2262: 2250: 2247: 2244: 2241: 2238: 2235: 2232: 2229: 2228: 2218: 2214: 2210: 2206: 2202: 2198: 2192: 2188: 2183: 2176: 2170: 2166: 2164: 2159: 2149: 2147: 2143: 2132: 2126: 2113: 2110: 2108: 2105: 2103: 2100: 2097: 2096: 2095: 2092: 2090: 2086: 2082: 2078: 2074: 2070: 2065: 2061: 2055: 2042: 2039: 2037: 2034: 2031: 2029: 2026: 2024: 2021: 2019: 2015: 2012: 2009: 2006: 2004: 2001: 1998: 1995: 1993: 1990: 1988: 1985: 1983: 1980: 1978: 1975: 1973: 1970: 1969: 1967: 1965: 1964:habitat types 1962:Other marine 1957: 1954: 1951: 1948: 1947: 1946: 1944: 1938: 1935: 1933: 1930: 1928: 1927:Littoral zone 1925: 1924: 1923: 1919: 1912:Marine biomes 1891: 1890: 1885: 1884: 1882: 1878: 1874: 1871: 1870: 1868: 1864: 1861: 1860: 1859: 1855: 1851: 1848: 1847: 1846: 1842: 1839: 1838: 1837: 1834: 1833: 1832: 1818: 1815: 1814: 1812: 1809: 1808: 1806: 1802: 1801: 1799: 1798: 1797: 1794: 1785: 1782: 1781: 1779: 1776: 1775: 1773: 1770: 1769: 1768: 1765: 1756: 1752: 1749: 1748: 1746: 1743: 1742: 1740: 1737: 1736: 1735: 1732: 1731: 1723: 1720: 1718: 1714: 1712: 1708: 1705: 1702: 1701: 1700: 1698: 1694: 1681: 1678: 1676: 1672: 1670: 1669:upland rivers 1666: 1664: 1660: 1658: 1654: 1652: 1651:upland rivers 1648: 1646: 1642: 1640: 1636: 1634: 1630: 1628: 1624: 1622: 1618: 1616: 1612: 1611: 1609: 1607: 1594: 1591: 1588: 1585: 1582: 1578: 1575: 1572: 1569: 1566: 1563: 1560: 1557: 1554: 1551: 1548: 1545: 1542: 1539: 1536: 1533: 1530: 1527: 1524: 1521: 1518: 1515: 1512: 1509: 1508: 1499: 1496: 1494: 1491: 1489: 1486: 1484: 1481: 1479: 1476: 1474: 1471: 1469: 1466: 1464: 1461: 1459: 1456: 1454: 1451: 1449: 1446: 1444: 1441: 1440: 1435: 1429: 1423: 1419: 1417: 1413: 1411: 1407: 1405: 1401: 1399: 1395: 1393: 1389: 1387: 1383: 1381: 1377: 1376: 1371: 1366: 1364: 1360: 1355: 1353: 1349: 1344: 1342: 1338: 1334: 1328: 1319: 1310: 1303: 1300: 1299: 1298: 1295: 1290: 1287: 1284: 1281: 1278: 1277: 1276: 1273: 1268: 1265: 1261: 1258: 1254: 1250: 1246: 1243: 1240: 1236: 1233: 1230: 1226: 1223: 1220: 1216: 1213: 1210: 1206: 1205: 1204: 1201: 1196: 1193: 1189: 1186: 1183: 1179: 1175: 1174: 1173: 1170: 1169: 1168: 1165: 1161: 1157: 1144: 1141: 1138: 1135: 1132: 1129: 1126: 1123: 1120: 1119: 1117: 1115: 1111: 1096: 1093: 1089: 1086: 1082: 1081: 1077: 1073: 1070: 1067: 1063: 1062: 1058: 1055: 1052: 1048: 1047: 1043: 1039: 1036: 1033: 1029: 1028: 1024: 1021: 1018: 1017: 1013: 1010: 1007: 1004: 1003:Mediterranean 1000: 999: 995: 992: 989: 985: 981: 980: 976: 972: 969: 966: 963: 959: 958: 955: 951: 948: 945: 941: 940: 936: 933: 930: 929: 926: 924: 919: 902: 898: 889: 885: 879: 876: 873: 870: 867: 863: 857: 854: 850: 849: 846: 837: 834: 831: 828: 825: 821: 815: 812: 809: 806: 803: 800: 797: 794: 791: 788: 785: 782: 779: 776: 773: 770: 769: 766: 765: 762: 761: 759: 757: 753: 739: 736: 734: 731: 729: 726: 724: 721: 719: 716: 714: 711: 709: 706: 704: 701: 699: 696: 694: 691: 689: 686: 684: 681: 679: 676: 672: 669: 667: 664: 662: 659: 657: 654: 652: 649: 648: 647: 644: 642: 639: 637: 634: 632: 629: 627: 624: 621: 617: 613: 610: 608: 605: 604: 603: 600: 598: 595: 593: 590: 588: 585: 581: 580:semideciduous 578: 576: 573: 572: 571: 568: 566: 563: 562: 555: 548: 545: 542: 539: 536: 535: 534: 528: 525: 522: 519: 518: 517: 515: 505: 503: 495: 492: 489: 486: 483: 480: 477: 476: 470: 467: 464: 456: 443: 440: 438: 434: 431: 429: 425: 423: 419: 416: 413: 412: 410: 406: 403: 401: 398: 396: 393: 391: 388: 386: 383: 381: 378: 376: 373: 371: 368: 366: 363: 362: 361: 358: 357: 356: 346: 343: 341: 338: 336: 335:High plateaus 333: 331: 328: 326: 323: 321: 318: 316: 313: 311: 308: 307: 306: 298: 295: 291: 287: 283: 277: 268: 259: 256: 252: 248: 244: 239: 230: 228: 224: 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Retrieved 4239: 4229: 4217:. Retrieved 4212: 4203: 4192:. Retrieved 4183: 4166: 4123: 4119: 4109: 4056: 4050: 4040: 3987: 3981: 3970: 3935: 3929: 3876:(1): 28234. 3873: 3867: 3857: 3812: 3806: 3796: 3751: 3745: 3697: 3693: 3683: 3630: 3624: 3614: 3563: 3559: 3549: 3498: 3494: 3483: 3473:, retrieved 3439: 3429: 3394: 3388: 3364:. Retrieved 3355: 3346: 3335:. Retrieved 3324: 3314: 3306:Google Books 3304:– via 3286: 3279: 3271:the original 3266: 3261: 3254: 3246:the original 3239: 3230: 3219:. Retrieved 3215:the original 3204: 3186: 3181: 3166:58:403–414, 3163: 3157: 3139: 3113:12:502–515, 3110: 3069: 3063: 3057: 3049:the original 3035: 3024: 3020: 3019:Schultz, J. 3015: 2992: 2988: 2982: 2959: 2955: 2949: 2941:the original 2931: 2924: 2913: 2908: 2898:. New York: 2895: 2856:(1): 1–239. 2853: 2847: 2841: 2828: 2822: 2814:the original 2808: 2801: 2791:. 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Index

Major habitat type
Biota

/ˈb.m/
geographical region
climate
vegetation
animal life
community
climate
ecosystems
habitats
microbiome
human microbiome
biosphere
Clements
biotic community
Möbius
phytophysiognomy
formation
vegetation
flora
species composition
Tansley
ecosystem
International Biological Program
Walter
biotope
biogeographic province
species composition

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