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Common toad

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1117:. Small, fast moving prey may be caught by a flick of the tongue while larger items are grabbed with the jaws. Having no teeth, it swallows food whole in a series of gulps. It does not recognise its prey as such but will try to consume any small, dark coloured, moving object it encounters at night. A research study showed that it would snap at a moving 1 cm (0.4 in) piece of black paper as if it were prey but would disregard a larger moving piece. Toads seem to use visual cues for feeding and can see their prey at low light intensities where humans are unable to discern anything. Periodically, the common toad sheds its skin. This comes away in tattered pieces and is then consumed. 4192: 1261: 1205: 1070: 933: 1385:
and they had reached 95% of their expected adult size. Some females that were on a biennial breeding cycle carried on growing rapidly for a longer time. Adjusting for differences in temperature and the length of the growing season, the toads grew and matured at much the same rate from the four colder localities. These juveniles reached maturity after 1.09 years for males and 1.55 years for females. However, the young toads from lowland France grew faster and longer to a much greater size taking an average 1.77 years for males and 2.49 years for females before reaching maturity.
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and hence of prowess. Nevertheless, fights occur in some instances. In a study at one pond where males outnumbered females by four or five to one, it was found that 38% of the males won the right to mate by defeating rivals in combat or by displacing other males already mounted on females. Male toads generally outnumber female toads at breeding ponds. A Swedish study found that female mortality was higher than that of males and that 41% of females did not come to the breeding pond in the spring and missed a year before reproducing again.
1239: 1680: 1821: 1724: 91: 1308:) but they are a darker colour, being blackish above and dark grey below. They can be distinguished from the tadpoles of other species by the fact that the mouth is the same width as the space between the eyes, and this is twice as large as the distance between the nostrils. Over the course of a few weeks their legs develop and their tail gradually gets reabsorbed. By twelve weeks of age they are miniature toads measuring about 1.5 cm (0.6 in) long and ready to leave the pond. 1552:, special tunnels have been constructed so that toads can cross under roads in safety. In other places, local wildlife groups run "toad patrols", carrying the amphibians across roads at busy crossing points in buckets. The toads start moving at dusk and for them to travel far, the temperature needs to remain above 5 °C (41 °F). On a warm wet night they may continue moving all night but if it cools down, they may stop earlier. An estimate was made of the significance of 643: 217: 1317: 31: 1446:". This is because it has a wide distribution and is, over most of its range, a common species. It is not particularly threatened by habitat loss because it is adaptable and is found in deciduous and coniferous forests, scrubland, meadows, parks and gardens. It prefers damp areas with dense foliage. The major threats it faces include loss of habitat locally, the drainage of wetlands where it breeds, agricultural activities, pollution, and mortality on roads. 1363: 66: 1281: 4150: 1520: 1325: 1292:. The males are enthusiastic, will try to grasp fish or inanimate objects and often mount the backs of other males. Sometimes several toads form a heap, each male trying to grasp the female at the base. It is a stressful period and mortality is high among breeding toads. A successful male stays in amplexus for several days and, as the female lays a long, double string of small black 1342:. This slows growth rates and reduces stamina and fitness. Larger juveniles at metamorphosis always outgrow smaller ones that have been reared in more crowded ponds. Even when they have heavy worm burdens, large juveniles grow faster than smaller individuals with light worm burdens. After several months of heavy worm infection, some juveniles in a study were only half as heavy as 1053:. The common toad is found at altitudes of up to 2,500 metres (8,200 ft) in the southern part of its range. It is largely found in forested areas with coniferous, deciduous and mixed woodland, especially in wet locations. It also inhabits open countryside, fields, copses, parks and gardens, and often occurs in dry areas well away from standing water. 1257:. They find their way to these by using a suite of orientation cues, including olfactory and magnetic cues, but also visual cues help guide their journeys. Toads experimentally moved elsewhere and fitted with tracking devices have been found to be able to locate their chosen breeding pond when the displacement exceeded three kilometres (two miles). 1384:
juveniles from different altitudes and latitudes, the specimens studied being from Norway, Germany, Switzerland, the Netherlands and France. At first the growth rates for males and females was identical. By the time they became mature their growth rate had slowed down to about 21% of the initial rate
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The common toad can reach about 15 cm (6 in) in length. Females are normally stouter than males and southern specimens tend to be larger than northern ones. The head is broad with a wide mouth below the terminal snout which has two small nostrils. There are no teeth. The bulbous, protruding
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The males arrive first and remain in the location for several weeks while the females only stay long enough to mate and spawn. Rather than fighting for the right to mate with a female, male toads may settle disputes by means of the pitch of their voice. Croaking provides a reliable sign of body size
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Many toads are killed by traffic while migrating to their breeding grounds. In Europe they have the highest rate of mortality from roadkill among amphibians. Many of the deaths take place on stretches of road where streams flow underneath showing that migration routes often follow water courses. In
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When attacked, the common toad adopts a characteristic stance, inflating its body and standing with its hindquarters raised and its head lowered. Its chief means of defence lies in the foul tasting secretion that is produced by its paratoid glands and other glands on its skin. This contains a toxin
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and reduced fitness as compared to nearby rural populations. The researchers demonstrated this by genetic analysis and by noting the greater number of physical abnormalities among urban as against rural tadpoles when raised in a controlled environment. It was considered that long term depletion in
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The toad has long been considered to be an animal of ill omen or a connection to a spirit world. This may have its origins in the fact that it is at home both on land and in the water. It may cause repugnance because of its drab, wart-like skin, its slow movements and the way it emerges from some
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in toad populations in the Netherlands. The number of females killed in the spring migration on a quiet country road (ten vehicles per hour) was compared with the number of strings of eggs laid in nearby fens. A 30% mortality rate was found, with the rate for deaths among males likely to be of a
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in spring and there is a mass migration towards the breeding sites. The toads converge on certain ponds that they favour while avoiding other stretches of water that seem eminently suitable. Adults use the same location year after year and over 80% of males marked as juveniles have been found to
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The common toad usually moves by walking rather slowly or in short shuffling jumps involving all four legs. It spends the day concealed in a lair that it has hollowed out under foliage or beneath a root or a stone where its colouring makes it inconspicuous. It emerges at dusk and may travel some
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tells how an old woman, a supposed witch, cursed her landlord and all his possessions when he demanded the unpaid rent for her cottage. Soon afterwards, a large toad fell on his wife and caused her to collapse. The toad was thrown into the fire but escaped with severe burns. Meanwhile, the old
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of various strengths. It was found that at certain concentrations, which were well above any normally found in the field, growth was increased and metamorphosis accelerated, but at others, there was no significant difference between the experimental tadpoles and controls. Nevertheless, certain
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on the first three fingers. He uses these to grasp the female when mating. The hind legs are short relative to other frogs' legs and the hind feet have long, unwebbed toes. There is no tail. The skin is dry and covered with small wart-like lumps. The colour is a fairly uniform shade of brown,
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Recuero E.; Canestrelli D.; Voeroes J.; Szabó, K.; Poyarkov, N. A.; Arntzen, J. W.; Crnobrnja-Isailovic, J.; Kidov A. A.; Cogălniceanu, D.; Caputo, F. P.; Nascetti, G.; Martínez-Solano, I. (2012). "Multilocus species tree analyses resolve the radiation of the widespread
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hatch out after 10 days. At first they cling to the remains of the strings and feed on the jelly. They later attach themselves to the underside of the leaves of water weed before becoming free swimming. The tadpoles at first look similar to those of the common frog
882:. The head joins the body without a noticeable neck and there is no external vocal sac. The body is broad and squat and positioned close to the ground. The fore limbs are short with the toes of the fore feet turning inwards. At breeding time, the male develops 921:
Common toads can live for many years and have survived for fifty years in captivity. In the wild, common toads are thought to live for about ten to twelve years. Their age can be determined by counting the number of annual growth rings in the bones of their
800:) seems to have arisen more recently. It is believed that the range of the ancestral form extended into Asia but that isolation between the eastern and western species complexes occurred as a result of the development of the Central Asian Deserts during the 1296:, he fertilises them with his sperm. As the pair wander piggyback around the shallow edges of the pond, the gelatinous egg strings, which may contain 1,500 to 6,000 eggs and be 3 to 4.5 metres (10 to 15 ft) in length, get tangled in plant stalks. 610:
Although toads are usually solitary animals, in the breeding season, large numbers of toads converge on certain breeding ponds, where the males compete to mate with the females. Eggs are laid in gelatinous strings in the water and later hatch out into
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over time as measured by sperm motility. This type of sperm senescence does not occur at a genetically fixed rate, but rather is influenced by environmental conditions that include availability of mating partners and temperature.
1475:. Both otter and frog seem to be extending their ranges to higher altitudes. The common toad cannot be legally sold or traded in the United Kingdom but there is a slow decline in toad numbers and it has therefore been declared a 1634:
contains enough similar toxin to cause serious symptoms (or even death) in animals, including humans. Clinical effects include severe irritation and pain to eyes, mouth, nose and throat, cardiovascular and respiratory symptoms,
630:. It is threatened by habitat loss, especially by drainage of its breeding sites, and some toads get killed on the roads as they make their annual migrations. It has long been associated in popular culture and literature with 1700:, it was thought to possess magical powers. Even ordinary people made use of dried toads, their bile, faeces and blood. In some areas, the finding of a toad in a house was considered evidence that a witch was present. In the 1230:) is unusual in that it can climb up water plants and move around on its muscular foot. It sometimes clings to the toe of a common toad and this is believed to be one of the means by which it disperses to new locations. 910:). The former is usually smaller and has a yellow band running down its back while the latter has a distinctive mottled pattern. The paratoid glands of both are parallel rather than slanting as in the common toad. The 1711:
investigated witchcraft in the region and searched the houses of suspected witches for dressed toads. He found none. These witches were reputed to use undomesticated toads as ingredients in their liniments and brews.
1224:, attacks adult common toads. It lays its eggs on the toad's skin and when these hatch, the larvae crawl into the toad's nostrils and eat its flesh internally with lethal consequences. The European fingernail clam ( 1350:
caused a decrease in food intake and some died. Another study investigated whether the use of nitrogenous fertilisers affects the development of common toad tadpoles. The toadlets were kept in dilute solutions of
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Dubois, Alain; Bour, Roger (2010). "The nomenclatural status of the nomina of amphibians and reptiles created by Garsault (1764), with a parsimonious solution to an old nomenclatural problem regarding the genus
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eyes have yellow or copper coloured irises and horizontal slit-shaped pupils. Just behind the eyes are two bulging regions, the paratoid glands, which are positioned obliquely. They contain a noxious substance,
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have been observed to puncture the skin with their beak and then peck out the toad's liver, thus avoiding the toxin. The tadpoles also exude noxious substances which deter fish from eating them but not the
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Garcia-Porta, J.; Litvinchuk, S. N.; Crochet, P. A.; Romano, A.; Lo-Valvo, M.; Lymberakis, P.; Carranza, S. (2012). "Molecular phylogenetics and historical biogeography of the west-palearctic common toads
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A General System of Nature,: Through the Three Grand Kingdoms of Animals, Vegetables, and Minerals, Systematically Divided Into Their Several Classes, Orders, Genera, Species, and Varieties (Volume 1)
41: 2148: 1128:, Scotland, observed a common toad moving along the bottom of the lake at a depth of 324 feet (99 m). They were surprised to find that an air-breathing animal could survive in such a location. 804:. The exact taxonomic relationships between these species remains unclear. A serological investigation into toad populations in Turkey undertaken in 2001 examined the blood serum proteins of 1742:
of the toad was considered poisonous and was known as "sweltered venom" and it was believed that it could spit or vomit poisonous fire. Toads were associated with devils and demons and in
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and his colleagues in 1922, and they succeeded in identifying its structure about 20 years later. Meanwhile, other researchers succeeded in isolating the same compound (and its parent
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distance in the dark while hunting. It is most active in wet weather. By morning it has returned to its base and may occupy the same place for several months. It is voracious and eats
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There are parts of its range where the common toad seems to be in decline. In Spain, increased aridity and habitat loss have led to a diminution in numbers and it is regarded as "
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Pettit, G. R.; Kamano, Y.; Drasar, P.; Inoue, M.; Knight, J. C. (1987). "Steroids and related natural products. 104. Bufadienolides. 36. Synthesis of bufalitoxin and bufotoxin".
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witch's cottage had caught fire and she was badly burnt. By next day, both toad and witch had died, and it was found that the woman's burns exactly mirrored those of the toad.
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Goater, Cameron P.; Semlitsch, Raymond D.; Bernasconi, Marco V. (1993). "Effects of body size and parasite infection on the locomotory performance of juvenile toads,
738:. It is found in France, the Iberian Peninsula and the Maghreb and grows to a larger size and has a spinier skin than its more northern counterparts with which it 1005:), the common toad is the fourth most common amphibian in Europe. It is found throughout the continent with the exception of Iceland, the cold northern parts of 837:
were being uplifted, an event which isolated the populations in the Iberian Peninsula from those in the rest of Europe. The remaining European lineage split into
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Agasyan, A.; Avisi, A.; Tuniyev, B.; Isailovic, J. C.; Lymberakis, P.; Andrén, C.; Cogalniceanu, D.; Wilkinson, J.; Ananjeva, N.; Üzüm, N.; et al. (2009).
3995: 1704:, the familiars were believed to be toads wearing elegant robes. These were herded by children who were being trained as witches. Between 1610 and 1612, the 1397:
in various holes in the ground, sometimes in basements, often in droves with other amphibians. Rarely they spend the winter in flowing waters with the
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Müller, L.; Hellmich, W. (1935). "Mitteilungenyüber die Herpetofauna der Iberischen Halbinsel. Über Salamandra salamandra almanzoris, n. ssp. und
4421: 772:, a group of closely related species which cannot be clearly demarcated. Several modern species are believed to form an ancient group of related 603:. The toad is an inconspicuous animal as it usually lies hidden during the day. It becomes active at dusk and spends the night hunting for the 713:
have been recognized over the years. The Caucasian toad is found in the mountainous regions of the Caucasus and was at one time classified as
4483: 4643: 812:. It found that the differences between the two were not significant and that therefore the former should be synonymized with the latter. 3281: 3256: 2684: 1651:. There is no known anti-venom. Treatment consists of supporting respiratory and cardiovascular functions, prevention of absorption and 1600:
had succeeded in synthesizing the toad toxin constituents bufotalin, bufalitoxin and bufotoxin. The chemical formula of bufotoxin is C
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at three to seven years old but there is great variability between populations. Juveniles are often parasitised by the lung nematode
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It has been found that urban populations of common toad occupying small areas and isolated by development show a lower level of
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on which it feeds. It moves with a slow, ungainly walk or short jumps, and has greyish-brown skin covered with wart-like lumps.
1696:, for whom a coat-of-arms was invented emblazoned with three toads. It was known that the toad could poison people and, as the 1362: 3575: 4653: 4607: 4117: 4088: 4012: 3978: 3948: 3916: 2827: 2662: 2638: 2457: 4439: 3222: 4658: 1121: 584: 4167: 891:
with the females being browner and the males greyer. The underside is a dirty white speckled with grey and black patches.
3968: 3938: 1620:, which, in small doses, increases the strength with which the heart muscle contracts; synthesized from foxglove plants ( 1439: 932: 627: 2053:
Specimen medicum, exhibens synopsin Reptilium emendatam cum experimentis circa venena et antidota Reptilium austriacorum
1811:", that when worn as a necklace or ring would warn the wearer of attempts to poison them. Shakespeare mentioned this in 815:
A study published in 2012 examined the phylogenetic relationships between the Eurasian and North African species in the
4576: 2334:(Anura, Bufonidae) populations in Southern Marmara (Manyas, Bahkesir) and Eastern Black Sea (Çamhhemşin, Rize) regions" 1693: 1288:
The males mount the females' backs, grasping them with their fore limbs under the armpits in a grip that is known as
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Xu, Q.; Oldham, R. S. (1997). "Lethal and sublethal effects of nitrogen fertilizer ammonium nitrate on common toad (
2072:(Amphibia, Anura), comments on the taxonomy of this genus, and comments on some nomina created by Laurenti (1768)". 4522: 4395: 2708: 2051: 1927: 887:
olive-brown or greyish-brown, sometimes partly blotched or banded with a darker shade. The common toad tends to be
2504: 2241: 2192: 4488: 4374: 1701: 918:) is also similar in appearance but it has a less rounded snout, damp smooth skin, and usually moves by leaping. 678: 619:
into tiny toads. The juveniles emerge from the water and remain largely terrestrial for the rest of their lives.
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Popular romances of the West of England; or, the drolls, traditions, and superstitions of Old Cornwall, Volume 2
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Santos, X.; Llorente, G. A.; Montori, A.; Carretero, M. A.; Franch, M.; Garriga, N.; Richter-Boix, A. (2007).
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Gelder, J. J. (1973). "A quantitative approach to the mortality resulting from traffic in a population of
4309: 4304: 2474: 1201:. These usually avoid the noxious secretion by puncturing the tadpole's skin and sucking out its juices. 2902:
Petkeviciute, R.; Stunzenas, V.; Staneviciute, G. (2004). "Cytogenetic and sequence comparison of adult
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described the emergence of the common toad from hibernation as one of the most moving signs of spring.
687: 4527: 1708: 1260: 1369: 595:. It is one of a group of closely related animals that are descended from a common ancestral line of 3650: 819:
group and indicated a long evolutionary history for the group. Nine to thirteen million years ago,
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and its colour tends to be blotched rather than uniform. It is now considered to be a synonym of
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numbers and habitat fragmentation can reduce population persistence in such urban environments.
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The common toad seems to be in decline in part of its range, but overall is listed as being of "
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from south Azerbaijan and Iran, split from the main lineage. Further divisions occurred with
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unusual swimming patterns and a few deformities were found among the experimental animals.
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Reading, C. J.; Loman, J.; Madsen, T. (1991). "Breeding pond fidelity in the common toad,
2906:(Digenea: Gorgoderidae) from the three-spined stickleback with larvae from two bivalves". 2566: 8: 4246: 4154: 3183:
Davies, N. B.; Halliday, T. R. (1979). "Competitive mate searching in male common toads,
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Ewart, J. P. (1987). "Neuroethology of releasing mechanisms: Prey-catching in toads".
2056:(in Latin). Viennae: Joan. Thom. Nob. de Trattnern. pp. i–ii + 1–215, plates 1–5. 4563: 4317: 4113: 4084: 3974: 3944: 3912: 3788: 3745: 3557: 3448: 3393: 3230: 3200: 3136:
Davies, N. B.; Halliday, T. R. (1978). "Deep croaks and fighting assessment in toads
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Larsen, Lis Olesen; Pedersen, Jan Nyholm (1981). "The snapping response of the toad,
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mountain range is facing predation by otters and increased competition from the frog
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Hemelaar, A. S. M.; van Gelder, J. J. (1979). "Annual growth rings in phalanges of
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Martens, R (1925). "Eine neue Eidechsengattung aus der Familie der Leposterniden".
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toad and in the book he composes a ditty in his own praise which starts like this:
1876: 1466: 1447: 1352: 1347: 1338: 1026: 751: 592: 4322: 4260: 4020: 4496: 3542: 3234: 2030: 1862: 1732: 1697: 1537: 1462: 1333: 1254: 895: 850: 769: 615:. After several months of growth and development, these sprout limbs and undergo 600: 4475: 3883: 2530: 2399: 2296: 1238: 1061: 4369: 4356: 4173: 4035: 3223:"Studies conducted at Lund University on amphibian research recently published" 2036: 1957: 1813: 1664: 1566: 1545: 1496: 1455: 821: 801: 781: 755: 739: 3673: 2919: 2735: 2350: 2329: 2085: 4663: 4637: 4447: 4231: 3792: 3784: 3474:
Hemelaar, Agnes (1988). "Age, growth and other population characteristics of
3054: 3005: 2901: 2591:(Anura, Amphibia) from the Netherlands and their use for age determination". 2026: 1922: 1820: 1744: 1648: 1541: 1500: 1484: 1443: 1381: 1293: 682:
in 1758. In this work, he placed all the frogs and toads in the single genus
673: 623: 616: 438: 383: 202: 75: 70: 4581: 2774: 2604: 1450:, an infectious disease of amphibians, has been reported in common toads in 1131:
The annual life cycle of the common toad is divided into three periods: the
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Chen, K. K.; Jensen, H.; Chen, A. L. (April 1932). "Action of Bufotoxins".
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L.) mit besonderer Berücksichtigung der Wanderung zu den Laichgewässern".
2822:(in Polish). Vol. 2: Płazy. Warszawa: Państwowe Wydawnictwo Naukowe. 1723: 4408: 4268: 4240: 1866: 1749: 1689: 1679: 1402: 1398: 1394: 1265: 1249: 1149: 1132: 1094: 1073: 1006: 998: 990: 982: 943: 911: 883: 846: 580: 216: 3838: 2408: 2305: 30: 4594: 4501: 4387: 4296: 3741: 3499: 3389: 3381: 3338: 3106: 2997: 2872: 2125: 1426: 1194: 894:
Other species with which the common toad could be confused include the
706: 642: 631: 588: 4413: 4273: 2756: 4343: 3518:"Post-meiotic intra-testicular sperm senescence in a wild vertebrate" 3161: 1808: 1807:
It was also believed that there was a jewel inside a toad's head, a "
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seem to be unaffected by it. Other predators of adult toads include
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Hettyey, A.; Vági, B.; Penn, D. J.; Hoi, H.; Wagner, R. H. (2012).
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Birstein, V. J.; Mazin, A. L. (1982). "Chromosomal polymorphism of
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in Siberia and its southerly range includes parts of northwestern
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Proceedings of the Society for Experimental Biology and Medicine
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Amphibian Species of the World: an Online Reference. Version 5.6
2221:, n. ssp., zwei neue Amphibienrassen aus der Sierra de Gredos". 2006:
Amphibian Species of the World: an Online Reference. Version 5.5
754:, a mountain range in central Spain. It has exceptionally large 4335: 2980:
Jungfer, Wolfgang (1943). "Beiträge zur Biologie der Erdkröte (
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gave instructions on using a toad in the concoction of spells:
1739: 1549: 1480: 1106: 1090: 1034: 1009:, Ireland and a number of Mediterranean islands. These include 939: 718: 112: 2565:. The Amphibian and Reptile Conservation Trust. Archived from 1519: 1299:
The strings of eggs absorb water and swell in size, and small
1753: 1451: 1418: 1324: 1169: 1114: 1050: 1014: 1010: 773: 561: 3182: 3135: 1049:. A closely related variant lives in eastern Asia including 2103: 1189:. Aquatic invertebrates that feed on toad tadpoles include 1181: 1173: 1161: 1086: 596: 568: 162: 142: 3582:. Amphibian and Reptile Conservation Trust. Archived from 4059: 4010: 2948: 2793: 1757: 1165: 1157: 1102: 1098: 3352:
Goater, C. P.; Ward, P. I. (1992). "Negative effects of
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Sinsch, Ulrich (1987). "Orientation behaviour of toads (
1865:. This has been dramatized by several authors including 849:. Very occasionally the common toad hybridizes with the 3695: 2761:, towards prey dummies at very low light intensities". 1380:
A comparison was made between the growth rate of newly
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found throughout most of Europe (with the exception of
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Wieland and Alles isolated bufotoxin from the skin of
3606:"Loss of genetic diversity and fitness in Common Toad 3425:
Archives of Environmental Contamination and Toxicology
2475:"Toads of the world: first, (some) toads of the north" 2216: 1855:
is one of the main characters in the children's novel
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Strijbosch, H. (1980). "Mortality in a population of
698:. The toads in this genus are included in the family 833:
splitting off about five million years ago when the
3418: 2447: 1499:, but during the 1990s, it became more abundant in 3970:Witch Hunts in Europe and America: An Encyclopedia 3940:Witch Hunts in Europe and America: An Encyclopedia 3515: 2986:Zeitschrift für Morphologie und Ökologie der Tiere 2146: 4042:. The museum of science, art and human perception 2066: 1483:, it is considered to be a "Rare Species" in the 776:from preglacial times. These are the spiny toad ( 4635: 3356:(Nematoda) on the growth and survival of toads ( 2528: 2239: 1999: 3909:Pagan Channel Islands: Europe's Hidden Heritage 3473: 3026: 2025: 1148:and is enough to deter many predators although 550:, or in Anglophone parts of Europe, simply the 3973:. Greenwood Publishing Group. pp. 20–21. 3511: 3509: 3088: 3084: 3082: 2842: 2499: 2497: 2495: 2493: 2491: 2039:. Lackington, Allen, and Co. pp. 648–649. 1639:and seizures, increased salivation, vomiting, 3932: 3930: 3928: 3715: 2660: 2452:. Harper Collins Publishers. pp. 73–74. 2450:Reptiles and Amphibians of Britain and Europe 845:less than three million years ago during the 4033: 3878: 3876: 3874: 3872: 3282:"Common toad (Bufo bufo) - British Reptiles" 3257:"Common toad (Bufo bufo) - British Reptiles" 2813: 2811: 2721: 2685:"Common toad (Bufo bufo) - British Reptiles" 2263: 1983:10.2305/IUCN.UK.2009.RLTS.T54596A11159939.en 1372:Tadpole development over a two-week period, 47:Common toad calls, recorded in Radnor, Wales 3604:Hitchings, S. P.; Beebee, T. J. C. (1998). 3506: 3079: 2631:The Observer's Book of British Wild Animals 2624: 2622: 2620: 2618: 2616: 2614: 2488: 2328:Tosunoğlua, Murat; Taskavak, Ertan (2001). 976: 3925: 2443: 2441: 2439: 2437: 2435: 2433: 2431: 2429: 2427: 2172: 2099: 2097: 2095: 1138: 1056: 215: 64: 29: 3943:. Greenwood Publishing Group. p. 7. 3869: 3625: 3610:populations isolated by inimical habitat" 3551: 3541: 3478:from different latitudes and altitudes". 3062: 3044: 3027:Landler, Lukas; Gollmann, Günter (2011). 2808: 2656: 2654: 2652: 2650: 2472: 2407: 2349: 2304: 1981: 1616:. Its physical effects resemble those of 1311: 1135:, the time of mating and feeding period. 665:The common toad was first given the name 3962: 3960: 2817: 2611: 2448:Arnold, Nicholas; Denys Ovenden (2002). 2049: 1995: 1993: 1842:Which, like the toad, ugly and venomous, 1819: 1722: 1678: 1674: 1518: 1323: 1315: 1279: 1259: 1237: 1203: 1076:eating adult common toad, Czech Republic 1068: 1060: 641: 38: 4103: 4078: 2979: 2424: 2092: 2043: 1969:The IUCN Red List of Threatened Species 1846:Wears yet a precious jewel in his head. 1626:), digoxin is used in the treatment of 1442:considers the common toad as being of " 1065:A common toad adopts a defensive stance 4636: 3906: 3698:. Toad watch: Helping toads to survive 2647: 2628: 2060: 1953: 1951: 1949: 1947: 1945: 1943: 1569:and skin of the common toad is called 1320:Tadpoles, some with hind legs, Germany 1253:return to the pond at which they were 4207: 4206: 3966: 3957: 3936: 3093:) displaced from the breeding site". 2633:. Frederick Warne. pp. 213–217. 2466: 2380:Molecular Phylogenetics and Evolution 2277:Molecular Phylogenetics and Evolution 2108:: Karyotype and C-banding pattern of 1990: 1756:as a toad when he poured poison into 4621:967CCA7F-43E7-40F9-B0BA-571DB3F20210 4375:77277517-b91f-4f54-878e-37c6ec7c7b56 3993: 3662:Animal Biodiversity and Conservation 2665:. Surrey Amphibian and Reptile Group 2330:"A serological investigation of the 2252:. American Museum of Natural History 2008:. American Museum of Natural History 1671:may prove useful in its management. 1523:Tunnel under road for toads, Germany 1122:remotely operated underwater vehicle 690:placed the common toad in the genus 4644:IUCN Red List least concern species 4036:"Frogs: Frog myths across cultures" 3627:10.1046/j.1420-9101.1998.11030269.x 3095:Journal of Comparative Physiology A 2789:"Experts' 'important' find in loch" 2378:species group (Anura, Bufonidae)". 1940: 1797:Boil thou first i' the charmed pot. 1692:, the toad was associated with the 1440:IUCN Red List of Threatened Species 1408: 1168:. Birds that feed on toads include 1037:in the northern mountain ranges of 878:, which is used to deter potential 734:. The spiny toad was classified as 628:IUCN Red List of Threatened Species 13: 4060:Shakespeare, William (1599–1600). 4011:Shakespeare, William (1605–1606). 2967:10.1111/j.1469-7998.1991.tb03811.x 1912:But none of them know half as much 1893:But never a name went down to fame 646:Skeleton showing shoulder girdle; 14: 4675: 4142: 3279: 3254: 2682: 2539:Reptiles and Amphibians of the UK 2147:Kuzmin, Sergius L. (2008-09-19). 1731:showing a witch feeding her toad 1630:. The skin of the South American 1458:and may affect some populations. 4190: 4148: 1928:Some Thoughts on the Common Toad 1885:The world has held great heroes, 1769: 1565:The main substance found in the 1361: 1029:. Its easterly range extends to 950: 931: 89: 4126: 4097: 4072: 4053: 4027: 4004: 3987: 3900: 3845: 3818: 3764: 3709: 3688: 3642: 3614:Journal of Evolutionary Biology 3597: 3568: 3467: 3412: 3345: 3298: 3273: 3248: 3215: 3176: 3129: 3020: 2973: 2942: 2895: 2836: 2781: 2750: 2715: 2701: 2676: 2580: 2551: 2522: 2366: 2321: 2240:Frost, Darrel R. (2013-01-09). 2233: 2210: 2000:Frost, Darrel R. (2011-01-31). 1908:Know all there is to be knowed. 1838:Sweet are the uses of adversity 1781:In the poison'd entrails throw. 1433: 1388: 1233: 679:10th edition of Systema Naturae 4168:BBC Nature on the Common Toad. 4136:Secker and Warburg 1950 p. 202 2818:Juszczyk, Włodzimierz (1987). 2593:Netherlands Journal of Zoology 2185: 2166: 2140: 2110:B. b. verrucosissima 2019: 1789:Days and nights has thirty-one 1493:Yamalo-Nenets Autonomous Okrug 868: 1: 2724:Behavioral and Brain Sciences 1934: 1889:As history books have showed; 1793:Swelter'd venom sleeping got, 1248:The common toad emerges from 1120:In 2007, researchers using a 796:). The European common toad ( 4654:Amphibians described in 1758 4066:, Act II, Scene I" 4017:, Act IV, Scene I" 3911:. Robert Hale. p. 118. 3827:Journal of Organic Chemistry 3543:10.1371/journal.pone.0050820 3233:. 2010-12-21. Archived from 3201:10.1016/0003-3472(79)90070-8 1777:Round about the cauldron go; 1688:dark hole. In Europe in the 1560: 591:, and in a small portion of 7: 4659:Taxa named by Carl Linnaeus 4019:. Poets.org. Archived from 2400:10.1016/j.ympev.2011.09.008 2297:10.1016/j.ympev.2011.12.019 1897:Compared with that of Toad! 1875:. Mr. Toad is a conceited, 1785:Toad, that under cold stone 1760:'s ear. The First Witch in 1573:. It was first isolated by 1514: 637: 10: 4680: 3967:Burns, William E. (2003). 3937:Burns, William E. (2003). 3857:Merriam-Webster Dictionary 2338:Italian Journal of Zoology 1655:to monitor the condition. 688:Josephus Nicolaus Laurenti 587:), in the western part of 4538: 4215: 4104:Grahame, Kenneth (1908). 4079:Grahame, Kenneth (1908). 3674:10.32800/abc.2007.30.0097 2920:10.1017/S0031182004006109 2736:10.1017/S0140525X00023128 2709:"AmphibiaWeb - Bufo bufo" 2529:Fairchild, G. J. (2003). 2351:10.1080/11250000109356402 2086:10.11646/zootaxa.2447.1.1 1346:. Their parasite-induced 843:Bufo verrucosissimus 806:Bufo verrucosissimus 672:by the Swedish biologist 299:Müller and Hellmich, 1935 235: 228: 224:Range map of common toad 223: 214: 191: 184: 86:Scientific classification 84: 62: 53: 37: 28: 23: 3785:10.3181/00379727-29-6141 3655:, near a breeding place" 2050:Laurenti, J. N. (1768). 1916:As intelligent Mr. Toad! 1904:The clever men at Oxford 1683:The Devil's coat of arms 1628:congestive heart failure 1598:Arizona State University 1596:By 1986, researchers at 1477:Biodiversity Action Plan 1465:". A population in the 1332:The common toad reaches 1208:Infected with larvae of 977:Distribution and habitat 742:. It is now accepted as 4110:The Wind in the Willows 4081:The Wind in the Willows 2847:resulting from the fly 2775:10.1163/156853882X00248 2605:10.1163/002829680X00069 1858:The Wind in the Willows 1709:Alonso de Salazar Frías 1139:Predators and parasites 1057:Behaviour and lifecycle 750:, is restricted to the 728:and is now accepted as 4000:. Hotten. p. 105. 3907:Peddle, S. V. (2007). 3480:Journal of Herpetology 2629:Stokeo, W. J. (1980). 1920: 1850: 1832: 1801: 1735: 1684: 1524: 1485:Bashkortostan Republic 1329: 1321: 1312:Development and growth 1285: 1273: 1245: 1215: 1077: 1066: 662: 521:Guérin-Méneville, 1838 470:Bufo vulgaris cinereus 256:Bufo (Bufo) grediscola 48: 4510:Paleobiology Database 4034:Wanner, Noel (2011). 3994:Hunt, Robert (1865). 3445:10.1007/s002449900188 3046:10.1186/1742-9994-8-6 2219:Bufo bufo gredosicola 1976:: e.T54596A11159939. 1881: 1834: 1823: 1773: 1726: 1682: 1675:Cultural significance 1522: 1479:priority species. In 1327: 1319: 1283: 1263: 1241: 1207: 1072: 1064: 995:Pelophylax esculentus 645: 585:Mediterranean islands 296:Bufo bufo gredosicola 259:Dubois and Bour, 2010 251:Dubois and Bour, 2010 46: 4649:Amphibians of Europe 4370:Fauna Europaea (new) 4162:Amphibians of Europe 4134:Shooting an Elephant 3033:Frontiers in Zoology 2820:Płazy i gady krajowe 2531:"Common Toad – 2151:Bufo verrucosissimus 1869:who called his play 1284:Egg strings, Belgium 1003:Lissotriton vulgaris 790:Japanese common toad 731:Bufo verrucosissimus 715:B. b. verrucosissima 518:Rana (Bufo) vulgaris 16:Species of amphibian 3839:10.1021/jo00392a014 3734:1973Oecol..13...93V 3534:2012PLoSO...750820H 3437:1997ArECT..32..298X 3374:1992Oecol..89..161G 3323:1993Oikos..66..129G 3154:1978Natur.274..683D 2865:1980Oecol..45..285S 2661:Daniel Winchester. 2392:2012MolPE..62...71R 2289:2012MolPE..63..113G 2275:species complex)". 1653:electrocardiography 904:European green toad 859:European green toad 822:Bufo eichwaldi 746:. The Gredos toad, 56:Conservation status 3742:10.1007/BF00379622 3382:10.1007/bf00317213 3107:10.1007/BF00605013 2998:10.1007/BF00421679 2955:Journal of Zoology 2873:10.1007/BF00346472 2849:Lucilia bufonivora 2559:"The common toad ( 2477:. Tetrapod zoology 2223:Zool. Anz. Leipzig 2126:10.1007/BF00133292 1833: 1827:as illustrated by 1736: 1706:Spanish inquisitor 1685: 1623:Digitalis purpurea 1525: 1489:Tatarstan Republic 1330: 1322: 1286: 1274: 1246: 1221:Lucilia bufonivora 1216: 1211:Lucilia bufonivora 1187:great crested newt 1078: 1067: 889:sexually dimorphic 831:Bufo spinosus 810:Bufo spinosus 786:B. verrucosissimus 760:Bufo spinosus 717:. It has a larger 702:, the true toads. 663: 49: 4631: 4630: 4209:Taxon identifiers 4119:978-0-674-03447-1 4090:978-0-674-03447-1 4040:The Exploratorium 3980:978-0-313-32142-9 3950:978-0-313-32142-9 3918:978-0-7090-8248-4 3833:(16): 3573–3578. 3280:Trust, Woodland. 3255:Trust, Woodland. 3231:HighBeam Research 3148:(5672): 683–685. 2829:978-83-01-05696-4 2797:. BBC. 2007-05-05 2763:Amphibia-Reptilia 2683:Trust, Woodland. 2640:978-0-7232-1503-5 2459:978-0-00-219964-3 1872:Toad of Toad Hall 1805: 1804: 1508:genetic diversity 1472:Pelophylax perezi 1417:intra-testicular 1344:control juveniles 1227:Sphaerium corneum 1218:A parasitic fly, 959: 827:described species 748:B. b. gredosicola 721:and differs from 599:and which form a 540: 539: 530: 522: 514: 506: 498: 486: 474: 466: 458: 450: 442: 435:Bufo tuberculosus 431: 423: 415: 403: 395: 387: 380:Bufo ferruginosus 376: 368: 360: 352: 344: 332: 320: 308: 300: 292: 284: 276: 268: 260: 252: 79: 44: 4671: 4624: 4623: 4611: 4610: 4598: 4597: 4585: 4584: 4572: 4571: 4559: 4558: 4557: 4531: 4530: 4518: 4517: 4505: 4504: 4492: 4491: 4479: 4478: 4476:NHMSYS0000080159 4466: 4465: 4456: 4455: 4443: 4442: 4430: 4429: 4417: 4416: 4404: 4403: 4391: 4390: 4378: 4377: 4365: 4364: 4352: 4351: 4339: 4338: 4326: 4325: 4313: 4312: 4300: 4299: 4290: 4289: 4277: 4276: 4264: 4263: 4251: 4250: 4249: 4236: 4235: 4234: 4204: 4203: 4195: 4194: 4193: 4186: 4153:Data related to 4152: 4137: 4132:Orwell, George 4130: 4124: 4123: 4101: 4095: 4094: 4076: 4070: 4069: 4057: 4051: 4050: 4048: 4047: 4031: 4025: 4024: 4008: 4002: 4001: 3991: 3985: 3984: 3964: 3955: 3954: 3934: 3923: 3922: 3904: 3898: 3897: 3895: 3894: 3880: 3867: 3866: 3864: 3863: 3849: 3843: 3842: 3822: 3816: 3815: 3768: 3762: 3761: 3713: 3707: 3706: 3704: 3703: 3692: 3686: 3685: 3659: 3646: 3640: 3639: 3629: 3601: 3595: 3594: 3592: 3591: 3572: 3566: 3565: 3555: 3545: 3513: 3504: 3503: 3471: 3465: 3464: 3416: 3410: 3409: 3354:Rhabdias bufonis 3349: 3343: 3342: 3302: 3296: 3295: 3293: 3292: 3277: 3271: 3270: 3268: 3267: 3252: 3246: 3245: 3243: 3242: 3219: 3213: 3212: 3195:(4): 1253–1267. 3189:Animal Behaviour 3180: 3174: 3173: 3162:10.1038/274683a0 3133: 3127: 3126: 3086: 3077: 3076: 3066: 3048: 3024: 3018: 3017: 2992:(1–3): 117–157. 2977: 2971: 2970: 2946: 2940: 2939: 2899: 2893: 2892: 2840: 2834: 2833: 2815: 2806: 2805: 2803: 2802: 2785: 2779: 2778: 2754: 2748: 2747: 2719: 2713: 2712: 2705: 2699: 2698: 2696: 2695: 2680: 2674: 2673: 2671: 2670: 2658: 2645: 2644: 2626: 2609: 2608: 2584: 2578: 2577: 2575: 2574: 2555: 2549: 2548: 2546: 2545: 2526: 2520: 2519: 2517: 2516: 2501: 2486: 2485: 2483: 2482: 2470: 2464: 2463: 2445: 2422: 2421: 2411: 2370: 2364: 2363: 2353: 2325: 2319: 2318: 2308: 2267: 2261: 2260: 2258: 2257: 2237: 2231: 2230: 2214: 2208: 2207: 2205: 2204: 2189: 2183: 2182: 2170: 2164: 2163: 2161: 2160: 2144: 2138: 2137: 2101: 2090: 2089: 2064: 2058: 2057: 2047: 2041: 2040: 2035:. Translated by 2023: 2017: 2016: 2014: 2013: 1997: 1988: 1987: 1985: 1955: 1770: 1698:witch's familiar 1575:Heinrich Wieland 1467:Sierra de Gredos 1448:Chytridiomycosis 1409:Sperm senescence 1365: 1353:ammonium nitrate 1339:Rhabdias bufonis 1328:Tadpole close-up 1027:Balearic Islands 961: 960: 935: 752:Sierra de Gredos 661: 657: 653: 649: 593:Northwest Africa 528: 520: 512: 504: 496: 484: 472: 464: 456: 454:Bufo ventricosus 449:Bonnaterre, 1789 448: 446:Bufo ventricosus 437: 429: 421: 413: 401: 393: 382: 375:Steenstrup, 1846 374: 366: 358: 350: 342: 330: 318: 306: 304:Bufo carbunculus 298: 290: 282: 274: 266: 258: 250: 248:Bufo (Bufo) bufo 219: 197: 94: 93: 73: 68: 67: 45: 33: 21: 20: 4679: 4678: 4674: 4673: 4672: 4670: 4669: 4668: 4634: 4633: 4632: 4627: 4619: 4614: 4606: 4601: 4593: 4588: 4580: 4575: 4567: 4562: 4553: 4552: 4547: 4534: 4526: 4521: 4513: 4508: 4500: 4497:Observation.org 4495: 4487: 4482: 4474: 4469: 4461: 4459: 4451: 4446: 4438: 4433: 4425: 4420: 4412: 4407: 4399: 4394: 4386: 4381: 4373: 4368: 4360: 4355: 4347: 4342: 4334: 4329: 4321: 4316: 4308: 4303: 4295: 4293: 4285: 4280: 4272: 4267: 4259: 4254: 4245: 4244: 4239: 4230: 4229: 4224: 4211: 4201: 4191: 4189: 4181: 4172:Information at 4166:Information at 4160:Information at 4145: 4140: 4131: 4127: 4120: 4102: 4098: 4091: 4077: 4073: 4058: 4054: 4045: 4043: 4032: 4028: 4009: 4005: 3992: 3988: 3981: 3965: 3958: 3951: 3935: 3926: 3919: 3905: 3901: 3892: 3890: 3882: 3881: 3870: 3861: 3859: 3851: 3850: 3846: 3823: 3819: 3769: 3765: 3714: 3710: 3701: 3699: 3694: 3693: 3689: 3657: 3647: 3643: 3602: 3598: 3589: 3587: 3574: 3573: 3569: 3514: 3507: 3492:10.2307/1564332 3472: 3468: 3417: 3413: 3350: 3346: 3331:10.2307/3545205 3303: 3299: 3290: 3288: 3278: 3274: 3265: 3263: 3253: 3249: 3240: 3238: 3221: 3220: 3216: 3181: 3177: 3134: 3130: 3087: 3080: 3025: 3021: 2978: 2974: 2947: 2943: 2900: 2896: 2841: 2837: 2830: 2816: 2809: 2800: 2798: 2787: 2786: 2782: 2755: 2751: 2720: 2716: 2707: 2706: 2702: 2693: 2691: 2681: 2677: 2668: 2666: 2659: 2648: 2641: 2627: 2612: 2585: 2581: 2572: 2570: 2557: 2556: 2552: 2543: 2541: 2527: 2523: 2514: 2512: 2503: 2502: 2489: 2480: 2478: 2473:Naish, Darren. 2471: 2467: 2460: 2446: 2425: 2371: 2367: 2326: 2322: 2268: 2264: 2255: 2253: 2238: 2234: 2215: 2211: 2202: 2200: 2191: 2190: 2186: 2175:Senckenbergiana 2171: 2167: 2158: 2156: 2145: 2141: 2102: 2093: 2065: 2061: 2048: 2044: 2027:von Linné, Carl 2024: 2020: 2011: 2009: 1998: 1991: 1956: 1941: 1937: 1919: 1914: 1910: 1906: 1900: 1895: 1891: 1887: 1877:anthropomorphic 1863:Kenneth Grahame 1849: 1844: 1840: 1800: 1795: 1791: 1787: 1783: 1779: 1677: 1615: 1611: 1607: 1603: 1563: 1557:similar order. 1538:the Netherlands 1528:some places in 1517: 1463:near threatened 1436: 1411: 1391: 1378: 1377: 1376: 1371: 1366: 1314: 1306:Rana temporaria 1236: 1141: 1113:and even small 1059: 987:Rana temporaria 979: 972: 962: 951: 946: 936: 916:Rana temporaria 896:natterjack toad 871: 851:natterjack toad 770:species complex 756:paratoid glands 726:morphologically 659: 655: 651: 648:1: suprascapula 647: 640: 601:species complex 536: 533: 422:Schneider, 1799 372:Bufo commutatus 351:Schneider, 1799 307:Bonaparte, 1835 275:Latreille, 1801 241: 240: 210: 199: 193: 180: 88: 80: 69: 65: 58: 39: 17: 12: 11: 5: 4677: 4667: 4666: 4661: 4656: 4651: 4646: 4629: 4628: 4626: 4625: 4612: 4599: 4586: 4573: 4560: 4544: 4542: 4536: 4535: 4533: 4532: 4519: 4506: 4493: 4480: 4467: 4457: 4444: 4431: 4418: 4405: 4392: 4379: 4366: 4357:Fauna Europaea 4353: 4340: 4327: 4314: 4301: 4291: 4278: 4265: 4252: 4237: 4221: 4219: 4213: 4212: 4200: 4199: 4177: 4176: 4174:Astolinto form 4170: 4164: 4158: 4157:at Wikispecies 4144: 4143:External links 4141: 4139: 4138: 4125: 4118: 4096: 4089: 4071: 4064:As you like it 4052: 4026: 4023:on 2007-01-10. 4003: 3986: 3979: 3956: 3949: 3924: 3917: 3899: 3868: 3844: 3817: 3763: 3708: 3687: 3641: 3620:(3): 269–283. 3596: 3576:"Common Toad: 3567: 3528:(12): e50820. 3505: 3486:(4): 369–388. 3466: 3431:(3): 298–303. 3411: 3368:(2): 161–165. 3344: 3317:(1): 129–136. 3297: 3286:Woodland Trust 3272: 3261:Woodland Trust 3247: 3227:Science Letter 3214: 3175: 3128: 3101:(5): 715–727. 3078: 3019: 2972: 2961:(2): 201–211. 2941: 2914:(6): 771–778. 2904:Phyllodistomum 2894: 2859:(2): 285–286. 2835: 2828: 2807: 2780: 2769:(4): 321–327. 2749: 2730:(3): 337–405. 2714: 2700: 2689:Woodland Trust 2675: 2646: 2639: 2610: 2599:(1): 129–135. 2579: 2550: 2521: 2509:: Common toad" 2487: 2465: 2458: 2423: 2365: 2344:(2): 165–168. 2320: 2283:(1): 113–130. 2262: 2232: 2209: 2184: 2165: 2139: 2091: 2059: 2042: 2037:William Turton 2018: 1989: 1938: 1936: 1933: 1901: 1882: 1835: 1814:As You Like It 1803: 1802: 1774: 1702:Basque Country 1676: 1673: 1665:cholestyramine 1649:hallucinations 1613: 1609: 1605: 1601: 1591:Bufo japonicus 1567:parotoid gland 1562: 1559: 1546:Northern Italy 1516: 1513: 1497:Irkutsk Oblast 1456:United Kingdom 1435: 1432: 1410: 1407: 1390: 1387: 1368: 1367: 1360: 1359: 1358: 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Anvari.org. 4067: 4065: 4056: 4041: 4037: 4030: 4022: 4018: 4016: 4007: 3999: 3998: 3990: 3982: 3976: 3972: 3971: 3963: 3961: 3952: 3946: 3942: 3941: 3933: 3931: 3929: 3920: 3914: 3910: 3903: 3889: 3888:Textfiles.com 3885: 3884:"Toad toxins" 3879: 3877: 3875: 3873: 3858: 3854: 3848: 3840: 3836: 3832: 3828: 3821: 3814: 3812: 3808: 3802: 3798: 3794: 3790: 3786: 3782: 3778: 3774: 3767: 3759: 3755: 3751: 3747: 3743: 3739: 3735: 3731: 3727: 3723: 3719: 3712: 3697: 3691: 3683: 3679: 3675: 3671: 3668:(1): 97–104. 3667: 3663: 3656: 3654: 3645: 3637: 3633: 3628: 3623: 3619: 3615: 3611: 3609: 3600: 3586:on 2011-09-05 3585: 3581: 3579: 3571: 3563: 3559: 3554: 3549: 3544: 3539: 3535: 3531: 3527: 3523: 3519: 3512: 3510: 3501: 3497: 3493: 3489: 3485: 3481: 3477: 3470: 3462: 3458: 3454: 3450: 3446: 3442: 3438: 3434: 3430: 3426: 3423:) tadpoles". 3422: 3415: 3407: 3403: 3399: 3395: 3391: 3387: 3383: 3379: 3375: 3371: 3367: 3363: 3359: 3355: 3348: 3340: 3336: 3332: 3328: 3324: 3320: 3316: 3312: 3308: 3301: 3287: 3283: 3276: 3262: 3258: 3251: 3237:on 2013-05-13 3236: 3232: 3228: 3224: 3218: 3210: 3206: 3202: 3198: 3194: 3190: 3186: 3179: 3171: 3167: 3163: 3159: 3155: 3151: 3147: 3143: 3139: 3132: 3124: 3120: 3116: 3112: 3108: 3104: 3100: 3096: 3092: 3085: 3083: 3074: 3070: 3065: 3060: 3056: 3052: 3047: 3042: 3038: 3034: 3030: 3023: 3015: 3011: 3007: 3003: 2999: 2995: 2991: 2988:(in German). 2987: 2983: 2976: 2968: 2964: 2960: 2956: 2952: 2945: 2937: 2933: 2929: 2925: 2921: 2917: 2913: 2909: 2905: 2898: 2890: 2886: 2882: 2878: 2874: 2870: 2866: 2862: 2858: 2854: 2850: 2846: 2839: 2831: 2825: 2821: 2814: 2812: 2796: 2795: 2790: 2784: 2776: 2772: 2768: 2764: 2760: 2753: 2745: 2741: 2737: 2733: 2729: 2725: 2718: 2710: 2704: 2690: 2686: 2679: 2664: 2663:"Common Toad" 2657: 2655: 2653: 2651: 2642: 2636: 2632: 2625: 2623: 2621: 2619: 2617: 2615: 2606: 2602: 2598: 2594: 2590: 2583: 2569:on 2008-11-21 2568: 2564: 2562: 2554: 2540: 2536: 2534: 2525: 2511:. 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Methuen. 4083:. Methuen. 3853:"Bufotoxin" 3807:B. vulgaris 3608:(Bufo bufo) 2002:"Bufonidae" 1867:A. A. Milne 1750:John Milton 1715:An English 1690:Middle Ages 1585:) from the 1403:green frogs 1266:mating ball 1250:hibernation 1164:, and even 1074:Grass snake 1007:Scandinavia 999:smooth newt 991:edible frog 983:common frog 944:Oxfordshire 912:common frog 869:Description 847:Pleistocene 778:B. spinosus 740:intergrades 656:3: clavicle 583:, and some 581:Scandinavia 579:, parts of 544:common toad 419:Bufo rubeta 407:Bufo rubeta 359:Bruch, 1862 291:Poche, 1912 4638:Categories 4555:Q109525973 4046:2012-08-23 3893:2012-05-26 3862:2012-05-26 3702:2012-04-30 3590:2012-05-04 3291:2023-03-21 3266:2023-03-21 3241:2012-04-18 2801:2014-07-31 2694:2023-03-11 2669:2012-05-03 2573:2012-05-04 2544:2012-03-16 2515:2012-05-04 2481:2012-06-23 2256:2013-01-24 2203:2012-09-17 2181:: 170–171. 2159:2012-09-17 2012:2012-08-24 1935:References 1495:, and the 1427:senescence 1425:undergoes 1268:(multiple 1124:to survey 1111:earthworms 997:) and the 981:After the 964:In a pond 938:Swimming, 902:) and the 788:) and the 707:subspecies 652:2: scapula 632:witchcraft 589:North Asia 343:Koch, 1872 331:Koch, 1872 319:Koch, 1872 4540:Rana bufo 4287:Bufo-bufo 4261:Bufo_bufo 4247:Bufo bufo 4217:Bufo bufo 4155:Bufo bufo 3793:1535-3699 3722:Oecologia 3718:Bufo bufo 3653:Bufo bufo 3578:Bufo bufo 3476:Bufo bufo 3421:Bufo bufo 3362:Oecologia 3358:Bufo bufo 3307:Bufo bufo 3185:Bufo bufo 3138:Bufo bufo 3091:Bufo bufo 3055:1742-9994 3006:0720-213X 2982:Bufo bufo 2951:Bufo bufo 2853:Oecologia 2845:Bufo bufo 2759:Bufo bufo 2744:144997714 2589:Bufo bufo 2561:Bufo bufo 2533:Bufo bufo 2507:Bufo bufo 2376:Bufo bufo 2332:Bufo bufo 2273:Bufo bufo 2106:Bufo bufo 1962:Bufo bufo 1809:toadstone 1752:depicted 1733:familiars 1717:folk tale 1669:lidocaine 1661:phenytoin 1637:paralysis 1632:cane toad 1583:bufotalin 1571:bufotoxin 1561:Bufotoxin 1191:dragonfly 1154:hedgehogs 1126:Loch Ness 924:phalanges 880:predators 876:bufotoxin 857:) or the 700:Bufonidae 696:Bufo bufo 626:" in the 557:Bufo bufo 526:Rana Bufo 411:robustior 195:Bufo bufo 171:Species: 153:Bufonidae 109:Kingdom: 103:Eukaryota 4549:Wikidata 4427:10193774 4294:BioLib: 4226:Wikidata 3801:87161465 3758:22706445 3750:28307986 3636:85117769 3562:23226542 3522:PLOS ONE 3461:35907368 3406:12236487 3398:28312868 3209:54259148 3123:26102029 3073:21418651 3039:(1): 6. 3014:46354366 2936:13313062 2928:15648700 2889:32817424 2881:28309542 2418:21964513 2360:83966131 2315:22214922 2229:: 49–57. 2134:37853177 2114:Genetica 2080:: 1–52. 2029:(1806). 1853:Mr. Toad 1825:Mr. Toad 1657:Atropine 1645:cyanosis 1554:roadkill 1515:Roadkill 1454:and the 1348:anorexia 1334:maturity 1301:tadpoles 1290:amplexus 1270:amplexus 1243:Amplexus 1214:, Poland 1193:larvae, 1083:woodlice 1025:and the 1023:Sardinia 966:Burgwald 835:Pyrenees 705:Various 638:Taxonomy 613:tadpoles 316:hybridus 230:Synonyms 203:Linnaeus 149:Family: 133:Amphibia 123:Chordata 119:Phylum: 113:Animalia 99:Domain: 76:IUCN 3.1 4616:ZooBank 4608:1103349 4595:8415029 4528:1350162 4388:5217160 4232:Q146375 4015:Macbeth 3730:Bibcode 3682:4570914 3553:3513296 3530:Bibcode 3500:1564332 3453:9096079 3433:Bibcode 3390:4219866 3370:Bibcode 3339:3545205 3319:Bibcode 3170:4222519 3150:Bibcode 3115:3119823 3064:3072941 2861:Bibcode 2388:Bibcode 2285:Bibcode 2074:Zootaxa 1765:Macbeth 1729:woodcut 1618:digoxin 1579:steroid 1534:Belgium 1530:Germany 1423:B. bufo 1374:Germany 1255:spawned 1146:bufagin 1144:called 1107:spiders 1091:beetles 1047:Tunisia 1043:Algeria 1039:Morocco 1031:Irkutsk 1019:Corsica 989:), the 970:Germany 780:), the 766:B. bufo 723:B. bufo 711:B. bufo 676:in the 577:Iceland 573:Ireland 560:, from 159:Genus: 139:Order: 129:Class: 74: ( 4515:387464 4440:173480 4414:326296 4362:177878 4349:BUFOBU 4336:333310 4310:101499 4183:Portal 4116:  4087:  3977:  3947:  3915:  3799:  3791:  3756:  3748:  3680:  3634:  3560:  3550:  3498:  3459:  3451:  3404:  3396:  3388:  3337:  3229:. via 3207:  3168:  3142:Nature 3121:  3113:  3071:  3061:  3053:  3012:  3004:  2934:  2926:  2887:  2879:  2826:  2742:  2637:  2456:  2416:  2358:  2313:  2132:  1831:, 1913 1740:saliva 1550:Poland 1491:, the 1487:, the 1481:Russia 1395:winter 1170:herons 1035:Africa 940:Otmoor 719:genome 441:, 1826 386:, 1826 328:medius 4582:10882 4577:EUNIS 4569:4RDKJ 4523:WoRMS 4460:NAS: 4453:54596 4422:IRMNG 4197:Frogs 3797:S2CID 3775:. 7. 3754:S2CID 3678:S2CID 3658:(PDF) 3632:S2CID 3496:JSTOR 3457:S2CID 3402:S2CID 3386:JSTOR 3335:JSTOR 3311:Oikos 3205:S2CID 3166:S2CID 3119:S2CID 3010:S2CID 2932:S2CID 2885:S2CID 2740:S2CID 2356:S2CID 2130:S2CID 1861:, by 1754:Satan 1727:1579 1694:Devil 1452:Spain 1419:sperm 1182:Crows 1174:crows 1099:flies 1087:slugs 1051:Japan 1015:Crete 1011:Malta 597:toads 562:Latin 492:var. 480:var. 439:Risso 409:var. 384:Risso 340:minor 326:var. 314:var. 143:Anura 4664:Bufo 4603:ITIS 4590:GBIF 4489:8384 4484:NCBI 4448:IUCN 4435:ITIS 4401:1952 4396:GISD 4383:GBIF 4344:EPPO 4323:NP2D 4305:BOLD 4114:ISBN 4085:ISBN 3975:ISBN 3945:ISBN 3913:ISBN 3789:ISSN 3746:PMID 3720:L". 3558:PMID 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Index


Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Amphibia
Anura
Bufonidae
Bufo
Binomial name
Linnaeus
1758

Synonyms
Risso
Risso
Latin
toad
Ireland
Iceland
Scandinavia
Mediterranean islands
North Asia
Northwest Africa
toads
species complex

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