909:
813:
861:
138:
845:
118:
893:
1475:
797:
829:
877:
966:
Hünefeld, Frank; Marvaldi, Adriana E.; Müller, Bianca; Lawrence, John F.; Beutel, Rolf Georg (2011). "The male postabdomen of the "ancestral" archostematan beetle
Tetraphalerus bruchi Heller, 1913 (Ommatidae) and its phylogenetic significance".
282:. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.
1184:
McKenna, Duane D.; Shin, Seunggwan; Ahrens, Dirk; Balke, Michael; Beza-Beza, Cristian; Clarke, Dave J.; Donath, Alexander; Escalona, Hermes E.; Friedrich, Frank; Letsch, Harald; Liu, Shanlin (2019-12-03).
244:
beetle family that has most ancestral characteristics. There are only seven extant species, confined to
Australia and South America. However, the geographical distribution was much wider during the
102:
1110:
937:
1056:
252:. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family:
3566:
3592:
1492:
1137:"Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea"
1118:
1020:
Yamamoto, Shûhei (2017). "A new genus of
Brochocoleini beetle in Upper Cretaceous Burmese amber (Coleoptera: Archostemata: Ommatidae)".
1429:
Li, Yan-Da; Tihelka, Erik; Yamamoto, Shûhei; Newton, Alfred F.; Xia, Fang-Yuan; Liu, Ye; Huang, Di-Ying; Cai, Chen-Yang (2023-08-22).
3416:
313:
3540:
941:
3579:
945:
1310:
3584:
1529:
908:
584:– Middle Triassic-Late Cretaceous (Eurasia) (other authors have considered this genus more closely related to Cupedidae)
1254:"The species-level phylogeny of archostematan beetles-where do Micromalthus debilis and Crowsoniella relicta belong?"
448:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian) (other authors consider this genus a junior synonym of
3597:
1389:(Coleoptera: Archostemata) from the Middle Jurassic Daohugou beds, with discussion on the generic circumscription"
218:
3661:
3506:
3501:
3478:
1253:
214:
137:
3656:
860:
3651:
1522:
812:
1335:
279:
3666:
3623:
3431:
892:
844:
3618:
3571:
1057:"Overview of and taphonomic bias in Archostemata (Insecta: Coleoptera) from Cretaceous amber"
3610:
3527:
2668:
1265:
1198:
1148:
1068:
1029:
976:
8:
3469:
2887:
1515:
1479:
1431:"Mesozoic Notocupes revealed as the sister group of Cupedidae (Coleoptera: Archostemata)"
1269:
1202:
1152:
1072:
1033:
980:
3646:
1365:
1281:
1229:
1186:
1092:
132:
1302:
Australian
Beetles Volume 2 : Archostemata, Myxophaga, Adephaga, Polyphaga (part)
3605:
3514:
3156:
2573:
1452:
1405:
1384:
1369:
1357:
1316:
1306:
1277:
1234:
1216:
1166:
1096:
1084:
1002:
724:– Early Jurassic-Early Cretaceous (Eurasia, ?Australia) (probably closely related to
1352:
1334:
Lee, Soo Bin; Nam, Gi Soo; Park, Jong Kyun; Lee, Byong Ho; Li, Yan-Da (2023-10-30).
1285:
796:
3253:
3032:
3027:
2950:
2708:
1724:
1442:
1400:
1347:
1273:
1224:
1206:
1156:
1076:
1037:
992:
984:
785:
was included in the family by some authors, but was excluded by
Kirejtshuk (2020).
1055:
Song, Zhenyu; Zhao, Xianye; Jarzembowski, Edmund A.; Xiao, Chuantao (2022-09-01).
1041:
3519:
3483:
2942:
2895:
2740:
2724:
2400:
2160:
2129:
2044:
1962:
1853:
1821:
705:
473:
1161:
1136:
828:
3345:
3296:
3151:
2958:
2626:
2610:
2549:
2444:
2415:
2343:
2307:
2291:
2207:
2152:
2057:
1954:
1803:
1765:
1745:
1641:
1080:
275:
248:
spanning across
Eurasia and Australia, suggesting that they were widespread on
1447:
1430:
1320:
988:
876:
3640:
3454:
3327:
3131:
3122:
2863:
2834:
2826:
2818:
2792:
2767:
2663:
2650:
2589:
2507:
2410:
2405:
2286:
2220:
2191:
2175:
2139:
2109:
2062:
2016:
1983:
1899:
1891:
1862:
1700:
1507:
1456:
1361:
1220:
1170:
1088:
867:
851:
750:
621:
361:
260:
59:
1211:
3304:
3228:
2974:
2966:
2921:
2905:
2855:
2776:
2732:
2684:
2602:
2515:
2384:
2371:
2351:
2312:
2212:
2199:
2170:
2104:
2092:
1988:
1945:
1928:
1920:
1870:
1708:
1678:
1609:
1598:
1582:
1238:
1006:
803:
589:
413:
341:
301:
241:
237:
199:
3493:
3553:
3463:
3390:
3361:
3335:
3322:
3262:
3220:
3204:
3196:
3188:
3172:
3164:
3100:
3058:
3050:
3019:
2934:
2913:
2879:
2871:
2784:
2716:
2700:
2692:
2655:
2642:
2618:
2536:
2528:
2478:
2470:
2428:
2392:
2363:
2335:
2330:
2246:
2241:
2233:
2183:
2119:
2114:
2028:
1993:
1978:
1970:
1886:
1878:
1695:
693:
661:
633:
513:
457:
429:
389:
377:
34:
1383:
Li, Y.-D.; Tihelka, E.; Newton, A. F.; Huang, D.-Y.; Cai, C.-Y. (2023).
117:
101:
3317:
3278:
3270:
3212:
3075:
3037:
2990:
2982:
2802:
2750:
2634:
2565:
2436:
2420:
2376:
2317:
2281:
2261:
2225:
2134:
2124:
1998:
1732:
1716:
997:
835:
778:
677:
565:
541:
485:
441:
369:
353:
79:
44:
3558:
1336:"Cretaceous beetles of the Jinju Formation (Coleoptera): Archostemata"
1300:
3382:
3374:
3366:
3353:
3309:
3108:
3083:
3011:
2839:
2810:
2494:
2325:
2299:
2251:
2165:
2067:
2036:
2008:
1841:
1816:
1811:
1792:
1757:
1629:
1621:
1570:
1558:
915:
899:
761:
717:
649:
605:
577:
497:
401:
271:
266:
169:
149:
84:
3545:
3425:
3448:
3286:
3236:
3180:
3139:
3088:
2745:
2676:
2557:
2269:
2075:
2049:
1907:
1740:
1687:
1666:
1576:
1485:
1474:
883:
819:
553:
525:
329:
274:
by some authors, but have been found to be more closely related to
245:
74:
69:
54:
49:
39:
3003:
2995:
2847:
2581:
2502:
2486:
89:
64:
3532:
757:– Middle Jurassic-Early Cretaceous (Asia) recent (South America)
2594:
2454:
2087:
1564:
1552:
1539:
365:
249:
233:
189:
179:
159:
965:
746:– Late Triassic-Late Cretaceous (Eurasia), recent (Australia)
700:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
656:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
628:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
560:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
508:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
492:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
480:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
424:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
408:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
384:– Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
739:
464:– Early Jurassic (Kyrgyzstan) (possibly closely related to
254:
668:– Late Triassic (Kyrgyzstan) (possibly closely related to
596:– Late Triassic (Kyrgyzstan) (possibly closely related to
290:
According to
Kirejtshuk, 2020 and subsequent literature.
1910:(furniture beetles, death watch beetles, spider beetles)
1054:
1428:
1183:
1187:"The evolution and genomic basis of beetle diversity"
1382:
3638:
2787:(jewel beetles, or metallic wood-boring beetles)
270:. The family is considered to be a subfamily of
1191:Proceedings of the National Academy of Sciences
2510:(true weevils, bark beetles, ambrosia beetles)
1677:
1537:
1523:
1333:
1251:
2753:(ironclad beetles, cylindrical bark beetles)
436:– Middle Jurassic-Early Cretaceous (Eurasia)
396:– Middle Jurassic-Early Cretaceous (Eurasia)
1530:
1516:
1134:
1117:. Texas A&M University. Archived from
572:– Middle Jurassic-Early Cretaceous (China)
348:– Early Jurassic-Late Cretaceous (Eurasia)
116:
100:
3417:List of subgroups of the order Coleoptera
2497:(straight snout weevils, New York weevil)
1596:
1446:
1404:
1351:
1228:
1210:
1160:
996:
2924:(cicada beetle, cicada parasite beetles)
1299:Adam., Slipinski (2020). "2:Ommatidae".
1019:
942:Department of the Environment and Energy
1135:Kirejtshuk, Alexander G. (2020-02-17).
308:– Early Jurassic-Late Cretaceous (Asia)
3639:
3014:(false fireflies, long-lipped beetles)
1839:
1790:
3430:
3429:
1838:
1789:
1664:
1663:
1595:
1511:
1424:
1422:
1420:
1418:
1416:
1298:
969:Arthropod Structure & Development
3040:(false metallic wood-boring beetles)
1130:
1128:
1252:Hörnschemeyer, Thomas (July 2009).
240:. The Ommatidae are considered the
13:
1435:Frontiers in Ecology and Evolution
1413:
14:
3678:
1467:
1125:
446:Kirejtshuk and Jarzembowski, 2020
358:Kirejtshuk and Jarzembowski, 2020
293:
2651:Perimylopidae, or Promecheilidae
2236:(minute brown scavenger beetles)
1473:
1406:10.11646/palaeoentomology.6.4.11
1278:10.1111/j.1365-3113.2009.00476.x
907:
891:
875:
859:
843:
827:
811:
795:
732:
324:– Early Cretaceous (South Korea)
136:
32:
2866:(variegated mud-loving beetles)
1376:
1353:10.11646/palaeoentomology.6.5.8
1327:
1292:
1245:
1177:
1103:
1048:
1013:
959:
930:
1:
2695:(narrow-waisted bark beetles)
2387:(parasitic flat bark beetles)
2338:(short-winged flower beetles)
2194:(lady beetles, or God's cows)
1931:(tooth-necked fungus beetles)
1115:Joel Hallan’s Biology Catalog
1061:Palaeontographica Abteilung A
1042:10.1016/j.cretres.2017.04.008
938:"Australian Faunal Directory"
923:
3207:(sand-loving scarab beetles)
2597:(minute tree-fungus beetles)
2423:(silvanid flat bark beetles)
548:– Late Triassic (Kyrgyzstan)
7:
3356:(primitive carrion beetles)
3312:(minute mud-loving beetles)
3175:(bumble bee scarab beetles)
2945:(soft-bodied plant beetles)
2916:(soft bodied plant beetles)
1719:(predaceous diving beetles)
1162:10.3390/geosciences10020073
698:Jarzembowski and Wang, 2016
688:– Late Jurassic (Australia)
364:, Myanmar, mid-Cretaceous (
336:– Middle Jurassic (England)
10:
3683:
3191:(scavenger scarab beetles)
3183:(enigmatic scarab beetles)
2356:Lamingtonium binnaberrense
2070:(soft-wing flower beetles)
788:
644:– Early Cretaceous (China)
616:– Early Cretaceous (China)
558:Li, Yamamoto and Cai, 2020
520:– Early Jurassic (England)
3438:
3414:
3344:
3330:(water scavenger beetles)
3295:
3261:
3252:
3130:
3121:
3074:
3063:Rhinorhipus tamborinensis
3049:
2933:
2904:
2842:(long-toed water beetles)
2801:
2775:
2766:
2735:(polypore fungus beetles)
2637:(palm and flower beetles)
2621:(tumbling flower beetles)
2568:(ant-like flower beetles)
2548:
2527:
2453:
2346:(lined flat bark beetles)
2320:(pleasing fungus beetles)
2260:
2215:(handsome fungus beetles)
2151:
2007:
1953:
1944:
1919:
1861:
1852:
1848:
1834:
1799:
1785:
1760:(burrowing water beetles)
1673:
1659:
1605:
1591:
1547:
1501:Atlas of Living Australia
1448:10.3389/fevo.2023.1015627
989:10.1016/j.asd.2010.12.003
536:– Middle Jurassic (China)
285:
133:Scientific classification
131:
124:
115:
108:
99:
23:
3377:(feather-winged beetles)
2711:(false longhorn beetles)
2658:(jugular-horned beetles)
2605:(false darkling beetles)
2576:(cryptic fungus beetles)
2395:(shining flower beetles)
1735:(crawling water beetles)
1634:Sikhotealinia zhiltzovae
1111:"Ommatidae Species List"
2858:(forest stream beetles)
2719:(synchroa bark beetles)
2679:(dead log bark beetles)
2671:(fire-coloured beetles)
2560:(ant-like leaf beetles)
2294:(silken fungus beetles)
2228:(well polished beetles)
2202:(minute fungus beetles)
2021:Acanthocnemus nigricans
1212:10.1073/pnas.1909655116
3369:(round fungus beetles)
2703:(false flower beetles)
2687:(wedge-shaped beetles)
2645:(false blister beetle)
2629:(hairy fungus beetles)
2584:(conifer bark beetles)
2481:(leaf-rolling weevils)
2142:(bark-gnawing beetles)
1902:(wounded-tree beetles)
1768:(false ground beetles)
1690:(trout-stream beetles)
1081:10.1127/pala/2022/0131
3619:Paleobiology Database
3281:(false clown beetles)
3103:(plate-thigh beetles)
2985:(false click beetles)
2890:(water-penny beetles)
2539:(ship-timber beetles)
2518:(pine flower weevils)
2366:(root-eating beetles)
2186:(minute bark beetles)
2097:Phloiophilus edwardsi
1624:(reticulated beetles)
1258:Systematic Entomology
916:Beutelius rutherfordi
280:molecular phylogenies
126:Tetraphalerus wagneri
3159:(false stag beetles)
3144:Belohina inexpectata
3006:(net-winged beetles)
2969:(rare click beetles)
2882:(travertine beetles)
2874:(minute mud beetles)
2439:(dry-fungus beetles)
2274:Agapytho foveicollis
2031:(false skin beetles)
1894:(Jacobson's beetles)
1614:Crowsoniella relicta
1305:. CSIRO Publishing.
772:– Recent (Australia)
3157:Diphyllostomatidae
2489:(primitive weevils)
2302:(flat bark beetles)
2052:(checkered beetles)
2039:(fruitworm beetles)
1727:(whirligig beetles)
1270:2009SysEn..34..533H
1203:2019PNAS..11624729M
1197:(49): 24729–24737.
1153:2020Geosc..10...73K
1073:2022PalAA.323..119S
1034:2017CrRes..76...34Y
1022:Cretaceous Research
981:2011ArtSD..40..146H
3662:Cretaceous insects
3022:(glowworm beetles)
2727:(darkling beetles)
2574:Archeocrypticidae
2431:(palmetto beetles)
2178:(dry bark beetles)
1965:(longhorn beetles)
783:Hong and Wang 1990
776:The extinct genus
712:– Eocene (England)
3634:
3633:
3606:Open Tree of Life
3432:Taxon identifiers
3423:
3422:
3410:
3409:
3406:
3405:
3402:
3401:
3394:
3386:
3385:(carrion beetles)
3378:
3370:
3357:
3331:
3313:
3282:
3274:
3248:
3247:
3240:
3232:
3224:
3216:
3208:
3200:
3192:
3184:
3176:
3168:
3160:
3147:
3117:
3116:
3104:
3096:
3066:
3041:
3023:
3015:
3007:
2999:
2986:
2978:
2970:
2962:
2961:(soldier beetles)
2954:
2951:Brachypsectridae
2946:
2925:
2917:
2891:
2883:
2875:
2867:
2859:
2851:
2843:
2830:
2822:
2814:
2788:
2762:
2761:
2754:
2736:
2728:
2720:
2712:
2709:Stenotrachelidae
2704:
2696:
2688:
2680:
2672:
2659:
2646:
2638:
2630:
2622:
2614:
2613:(blister beetles)
2606:
2598:
2585:
2577:
2569:
2561:
2540:
2519:
2511:
2498:
2490:
2482:
2474:
2464:
2440:
2432:
2424:
2396:
2388:
2380:
2367:
2359:
2347:
2339:
2321:
2303:
2295:
2277:
2237:
2229:
2216:
2203:
2195:
2187:
2179:
2161:Akalyptoischiidae
2143:
2100:
2083:
2071:
2053:
2040:
2032:
2024:
1974:
1966:
1940:
1939:
1932:
1911:
1903:
1895:
1882:
1874:
1830:
1829:
1807:
1781:
1780:
1777:
1776:
1769:
1761:
1753:
1736:
1728:
1720:
1712:
1704:
1691:
1655:
1654:
1637:
1625:
1617:
1312:978-0-643-09731-5
900:Beutelius sagitta
784:
771:
756:
745:
723:
711:
699:
687:
667:
666:Ponomarenko, 1966
655:
643:
627:
615:
595:
594:Ponomarenko, 1966
583:
582:Ponomarenko, 1964
571:
559:
547:
546:Ponomarenko, 1966
535:
530:Li and Cai in Li
519:
507:
491:
479:
463:
462:Ponomarenko, 1969
447:
435:
423:
407:
395:
383:
359:
347:
335:
323:
307:
226:
225:
222:
16:Family of beetles
3674:
3657:Jurassic insects
3627:
3626:
3614:
3613:
3601:
3600:
3588:
3587:
3575:
3574:
3562:
3561:
3549:
3548:
3536:
3535:
3523:
3522:
3510:
3509:
3497:
3496:
3487:
3486:
3474:
3473:
3472:
3459:
3458:
3457:
3427:
3426:
3392:
3384:
3376:
3368:
3355:
3329:
3311:
3280:
3272:
3259:
3258:
3254:Staphyliniformia
3238:
3230:
3222:
3214:
3206:
3198:
3190:
3182:
3174:
3166:
3158:
3141:
3128:
3127:
3102:
3093:Declinia relicta
3090:
3060:
3039:
3033:Sinopyrophoridae
3028:Rhagophthalmidae
3021:
3013:
3005:
2997:
2984:
2976:
2968:
2960:
2952:
2944:
2943:Artematopodidae
2923:
2915:
2889:
2881:
2873:
2865:
2857:
2850:(riffle beetles)
2849:
2841:
2829:(turtle beetles)
2828:
2820:
2812:
2786:
2773:
2772:
2752:
2734:
2726:
2718:
2710:
2702:
2694:
2686:
2678:
2670:
2657:
2644:
2636:
2628:
2620:
2612:
2604:
2596:
2583:
2575:
2567:
2559:
2538:
2517:
2509:
2496:
2488:
2480:
2473:(fungus weevils)
2472:
2463:
2462:
2458:
2445:Tasmosalpingidae
2438:
2430:
2422:
2394:
2386:
2378:
2365:
2353:
2345:
2337:
2319:
2301:
2293:
2271:
2235:
2227:
2214:
2201:
2193:
2185:
2177:
2141:
2094:
2077:
2069:
2051:
2038:
2030:
2018:
2017:Acanthocnemidae
1972:
1964:
1951:
1950:
1930:
1909:
1901:
1893:
1880:
1872:
1859:
1858:
1850:
1849:
1836:
1835:
1805:
1787:
1786:
1767:
1759:
1747:
1734:
1726:
1718:
1710:
1703:(ground beetles)
1702:
1689:
1675:
1674:
1661:
1660:
1631:
1623:
1611:
1610:Crowsoniellidae
1593:
1592:
1532:
1525:
1518:
1509:
1508:
1504:
1478:Data related to
1477:
1461:
1460:
1450:
1426:
1411:
1410:
1408:
1393:Palaeoentomology
1385:"New species of
1380:
1374:
1373:
1355:
1340:Palaeoentomology
1331:
1325:
1324:
1296:
1290:
1289:
1249:
1243:
1242:
1232:
1214:
1181:
1175:
1174:
1164:
1132:
1123:
1122:
1107:
1101:
1100:
1067:(4–6): 119–137.
1052:
1046:
1045:
1017:
1011:
1010:
1000:
963:
957:
956:
954:
953:
944:. Archived from
934:
911:
895:
879:
863:
847:
831:
815:
799:
782:
765:
755:Waterhouse, 1901
754:
743:
722:Handlirsch, 1906
721:
709:
697:
681:
665:
654:Kirejtshuk, 2020
653:
637:
625:
609:
593:
581:
569:
557:
545:
529:
517:
501:
490:Li and Cai, 2021
489:
478:Kirejtshuk, 2020
477:
461:
445:
433:
417:
406:Kirejtshuk, 2020
405:
393:
382:Kirejtshuk, 2020
381:
357:
345:
333:
317:
306:Kirejtshuk, 2020
305:
236:in the suborder
232:are a family of
213:
141:
140:
120:
104:
94:
31:
27:Temporal range:
21:
20:
3682:
3681:
3677:
3676:
3675:
3673:
3672:
3671:
3652:Beetle families
3637:
3636:
3635:
3630:
3622:
3617:
3609:
3604:
3596:
3591:
3583:
3578:
3570:
3565:
3557:
3552:
3544:
3539:
3531:
3526:
3518:
3513:
3505:
3500:
3492:
3490:
3482:
3477:
3468:
3467:
3462:
3453:
3452:
3447:
3434:
3424:
3419:
3398:
3340:
3291:
3273:(clown beetles)
3244:
3215:(betsy beetles)
3123:Scarabaeiformia
3113:
3070:
3045:
2977:(click beetles)
2953:(Texas beetles)
2929:
2900:
2896:Ptilodactylidae
2821:(cedar beetles)
2797:
2758:
2741:Trictenotomidae
2627:Mycetophagidae
2544:
2523:
2460:
2459:
2457:
2449:
2401:Phloeostichidae
2352:Lamingtoniidae
2344:Laemophloeidae
2292:Cryptophagidae
2256:
2147:
2130:Thanerocleridae
2093:Phloiophilidae
2080:Metaxina ornata
2045:Chaetosomatidae
2003:
1936:
1915:
1873:(auger beetles)
1854:Bostrichiformia
1844:
1826:
1822:Torridincolidae
1806:(skiff beetles)
1804:Hydroscaphidae
1795:
1773:
1766:Trachypachidae
1711:(tiger beetles)
1669:
1651:
1601:
1587:
1543:
1536:
1491:
1470:
1465:
1464:
1427:
1414:
1381:
1377:
1332:
1328:
1313:
1297:
1293:
1250:
1246:
1182:
1178:
1133:
1126:
1109:
1108:
1104:
1053:
1049:
1018:
1014:
964:
960:
951:
949:
936:
935:
931:
926:
919:
912:
903:
896:
887:
880:
871:
864:
855:
848:
839:
832:
823:
816:
807:
800:
791:
735:
706:Tetraphalerites
474:Jarzembowskiops
296:
288:
212:
135:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
30:Ladinian–Recent
29:
28:
25:
17:
12:
11:
5:
3680:
3670:
3669:
3664:
3659:
3654:
3649:
3632:
3631:
3629:
3628:
3615:
3602:
3589:
3576:
3563:
3550:
3537:
3524:
3511:
3498:
3488:
3475:
3460:
3444:
3442:
3436:
3435:
3421:
3420:
3415:
3412:
3411:
3408:
3407:
3404:
3403:
3400:
3399:
3397:
3396:
3393:(rove beetles)
3391:Staphylinidae
3388:
3380:
3372:
3364:
3359:
3350:
3348:
3346:Staphylinoidea
3342:
3341:
3339:
3338:
3333:
3328:Hydrophilidae
3325:
3320:
3315:
3307:
3301:
3299:
3297:Hydrophiloidea
3293:
3292:
3290:
3289:
3284:
3276:
3267:
3265:
3256:
3250:
3249:
3246:
3245:
3243:
3242:
3239:(hide beetles)
3234:
3226:
3223:(rain beetles)
3218:
3210:
3202:
3199:(stag beetles)
3194:
3186:
3178:
3170:
3162:
3154:
3152:Bolboceratidae
3149:
3136:
3134:
3125:
3119:
3118:
3115:
3114:
3112:
3111:
3106:
3098:
3086:
3080:
3078:
3072:
3071:
3069:
3068:
3059:Rhinorhipidae
3055:
3053:
3051:Rhinorhipoidea
3047:
3046:
3044:
3043:
3035:
3030:
3025:
3017:
3009:
3001:
2993:
2988:
2980:
2972:
2964:
2956:
2948:
2939:
2937:
2931:
2930:
2928:
2927:
2919:
2910:
2908:
2902:
2901:
2899:
2898:
2893:
2885:
2877:
2869:
2864:Heteroceridae
2861:
2853:
2845:
2837:
2832:
2827:Chelonariidae
2824:
2819:Callirhipidae
2816:
2813:(pill beetles)
2807:
2805:
2799:
2798:
2796:
2795:
2790:
2781:
2779:
2770:
2764:
2763:
2760:
2759:
2757:
2756:
2748:
2743:
2738:
2730:
2725:Tenebrionidae
2722:
2714:
2706:
2698:
2690:
2682:
2674:
2666:
2661:
2653:
2648:
2640:
2632:
2624:
2616:
2608:
2600:
2592:
2587:
2579:
2571:
2563:
2554:
2552:
2550:Tenebrionoidea
2546:
2545:
2543:
2542:
2533:
2531:
2525:
2524:
2522:
2521:
2513:
2508:Curculionidae
2505:
2500:
2492:
2484:
2476:
2467:
2465:
2455:Curculionoidea
2451:
2450:
2448:
2447:
2442:
2434:
2426:
2418:
2416:Protocucujidae
2413:
2408:
2403:
2398:
2390:
2382:
2374:
2369:
2361:
2349:
2341:
2333:
2328:
2323:
2315:
2310:
2308:Cybocephalidae
2305:
2297:
2289:
2284:
2279:
2266:
2264:
2258:
2257:
2255:
2254:
2249:
2244:
2239:
2231:
2223:
2218:
2210:
2208:Discolomatidae
2205:
2197:
2192:Coccinellidae
2189:
2181:
2176:Bothrideridae
2173:
2168:
2163:
2157:
2155:
2153:Coccinelloidea
2149:
2148:
2146:
2145:
2140:Trogossitidae
2137:
2132:
2127:
2122:
2117:
2112:
2107:
2102:
2090:
2085:
2073:
2065:
2060:
2058:Lophocateridae
2055:
2047:
2042:
2034:
2026:
2013:
2011:
2005:
2004:
2002:
2001:
1996:
1991:
1986:
1981:
1976:
1973:(leaf beetles)
1971:Chrysomelidae
1968:
1959:
1957:
1955:Chrysomeloidea
1948:
1942:
1941:
1938:
1937:
1935:
1934:
1925:
1923:
1917:
1916:
1914:
1913:
1905:
1900:Nosodendridae
1897:
1892:Jacobsoniidae
1889:
1884:
1881:(skin beetles)
1876:
1867:
1865:
1856:
1846:
1845:
1832:
1831:
1828:
1827:
1825:
1824:
1819:
1814:
1809:
1800:
1797:
1796:
1783:
1782:
1779:
1778:
1775:
1774:
1772:
1771:
1763:
1755:
1750:Meru phyllisae
1743:
1738:
1730:
1722:
1714:
1706:
1698:
1693:
1684:
1682:
1671:
1670:
1657:
1656:
1653:
1652:
1650:
1649:
1644:
1642:Micromalthidae
1639:
1627:
1619:
1606:
1603:
1602:
1589:
1588:
1586:
1585:
1579:
1573:
1567:
1561:
1555:
1548:
1545:
1544:
1535:
1534:
1527:
1520:
1512:
1506:
1505:
1489:
1483:
1482:at Wikispecies
1469:
1468:External links
1466:
1463:
1462:
1412:
1399:(4): 398–415.
1375:
1326:
1311:
1291:
1264:(3): 533–558.
1244:
1176:
1124:
1121:on 2012-10-23.
1102:
1047:
1012:
975:(2): 146–158.
958:
928:
927:
925:
922:
921:
920:
913:
906:
904:
897:
890:
888:
881:
874:
872:
865:
858:
856:
849:
842:
840:
833:
826:
824:
817:
810:
808:
801:
794:
790:
787:
774:
773:
758:
747:
734:
731:
730:
729:
713:
701:
689:
673:
657:
645:
629:
626:Yamamoto, 2017
617:
601:
585:
573:
561:
549:
537:
521:
509:
493:
481:
469:
453:
437:
425:
409:
397:
385:
373:
349:
337:
334:Westwood, 1854
325:
309:
295:
294:Extinct genera
292:
287:
284:
276:Micromalthidae
224:
223:
207:
203:
202:
197:
193:
192:
187:
183:
182:
177:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
129:
128:
122:
121:
113:
112:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
3679:
3668:
3667:Mesozoic Asia
3665:
3663:
3660:
3658:
3655:
3653:
3650:
3648:
3645:
3644:
3642:
3625:
3620:
3616:
3612:
3607:
3603:
3599:
3594:
3590:
3586:
3581:
3577:
3573:
3568:
3564:
3560:
3555:
3551:
3547:
3542:
3538:
3534:
3529:
3525:
3521:
3516:
3512:
3508:
3503:
3499:
3495:
3489:
3485:
3480:
3476:
3471:
3465:
3461:
3456:
3450:
3446:
3445:
3443:
3441:
3437:
3433:
3428:
3418:
3413:
3395:
3389:
3387:
3381:
3379:
3373:
3371:
3365:
3363:
3360:
3358:
3352:
3351:
3349:
3347:
3343:
3337:
3334:
3332:
3326:
3324:
3321:
3319:
3316:
3314:
3308:
3306:
3303:
3302:
3300:
3298:
3294:
3288:
3285:
3283:
3279:Sphaeritidae
3277:
3275:
3269:
3268:
3266:
3264:
3260:
3257:
3255:
3251:
3241:
3235:
3233:
3229:Scarabaeidae
3227:
3225:
3219:
3217:
3211:
3209:
3203:
3201:
3195:
3193:
3187:
3185:
3179:
3177:
3171:
3169:
3167:(dor beetles)
3163:
3161:
3155:
3153:
3150:
3148:
3145:
3138:
3137:
3135:
3133:
3132:Scarabaeoidea
3129:
3126:
3124:
3120:
3110:
3107:
3105:
3099:
3097:
3094:
3087:
3085:
3082:
3081:
3079:
3077:
3073:
3067:
3064:
3057:
3056:
3054:
3052:
3048:
3042:
3036:
3034:
3031:
3029:
3026:
3024:
3018:
3016:
3010:
3008:
3002:
3000:
2994:
2992:
2989:
2987:
2981:
2979:
2973:
2971:
2967:Cerophytidae
2965:
2963:
2957:
2955:
2949:
2947:
2941:
2940:
2938:
2936:
2932:
2926:
2922:Rhipiceridae
2920:
2918:
2912:
2911:
2909:
2907:
2903:
2897:
2894:
2892:
2886:
2884:
2878:
2876:
2870:
2868:
2862:
2860:
2856:Eulichadidae
2854:
2852:
2846:
2844:
2838:
2836:
2835:Cneoglossidae
2833:
2831:
2825:
2823:
2817:
2815:
2809:
2808:
2806:
2804:
2800:
2794:
2793:Schizopodidae
2791:
2789:
2783:
2782:
2780:
2778:
2774:
2771:
2769:
2768:Elateriformia
2765:
2755:
2749:
2747:
2744:
2742:
2739:
2737:
2733:Tetratomidae
2731:
2729:
2723:
2721:
2715:
2713:
2707:
2705:
2699:
2697:
2691:
2689:
2685:Ripiphoridae
2683:
2681:
2675:
2673:
2669:Pyrochroidae
2667:
2665:
2664:Pterogeniidae
2662:
2660:
2654:
2652:
2649:
2647:
2641:
2639:
2633:
2631:
2625:
2623:
2617:
2615:
2609:
2607:
2603:Melandryidae
2601:
2599:
2593:
2591:
2590:Chalcodryidae
2588:
2586:
2580:
2578:
2572:
2570:
2564:
2562:
2556:
2555:
2553:
2551:
2547:
2541:
2535:
2534:
2532:
2530:
2526:
2520:
2516:Nemonychidae
2514:
2512:
2506:
2504:
2501:
2499:
2493:
2491:
2485:
2483:
2477:
2475:
2469:
2468:
2466:
2456:
2452:
2446:
2443:
2441:
2435:
2433:
2427:
2425:
2419:
2417:
2414:
2412:
2411:Propalticidae
2409:
2407:
2406:Priasilphidae
2404:
2402:
2399:
2397:
2391:
2389:
2385:Passandridae
2383:
2381:
2379:(sap beetles)
2375:
2373:
2370:
2368:
2362:
2360:
2357:
2350:
2348:
2342:
2340:
2334:
2332:
2329:
2327:
2324:
2322:
2316:
2314:
2311:
2309:
2306:
2304:
2298:
2296:
2290:
2288:
2287:Cavognathidae
2285:
2283:
2280:
2278:
2275:
2268:
2267:
2265:
2263:
2259:
2253:
2250:
2248:
2245:
2243:
2240:
2238:
2232:
2230:
2224:
2222:
2221:Eupsilobiidae
2219:
2217:
2213:Endomychidae
2211:
2209:
2206:
2204:
2200:Corylophidae
2198:
2196:
2190:
2188:
2182:
2180:
2174:
2172:
2169:
2167:
2164:
2162:
2159:
2158:
2156:
2154:
2150:
2144:
2138:
2136:
2133:
2131:
2128:
2126:
2123:
2121:
2118:
2116:
2115:Protopeltidae
2113:
2111:
2110:Prionoceridae
2108:
2106:
2103:
2101:
2098:
2091:
2089:
2086:
2084:
2081:
2074:
2072:
2066:
2064:
2063:Mauroniscidae
2061:
2059:
2056:
2054:
2048:
2046:
2043:
2041:
2035:
2033:
2027:
2025:
2022:
2015:
2014:
2012:
2010:
2006:
2000:
1997:
1995:
1992:
1990:
1987:
1985:
1984:Megalopodidae
1982:
1980:
1977:
1975:
1969:
1967:
1963:Cerambycidae
1961:
1960:
1958:
1956:
1952:
1949:
1947:
1943:
1933:
1929:Derodontidae
1927:
1926:
1924:
1922:
1921:Derodontoidea
1918:
1912:
1906:
1904:
1898:
1896:
1890:
1888:
1887:Endecatomidae
1885:
1883:
1877:
1875:
1871:Bostrichidae
1869:
1868:
1866:
1864:
1863:Bostrichoidea
1860:
1857:
1855:
1851:
1847:
1843:
1837:
1833:
1823:
1820:
1818:
1817:Sphaeriusidae
1815:
1813:
1810:
1808:
1802:
1801:
1798:
1794:
1788:
1784:
1770:
1764:
1762:
1756:
1754:
1751:
1744:
1742:
1739:
1737:
1731:
1729:
1723:
1721:
1715:
1713:
1709:Cicindelidae
1707:
1705:
1699:
1697:
1694:
1692:
1686:
1685:
1683:
1680:
1676:
1672:
1668:
1662:
1658:
1648:
1645:
1643:
1640:
1638:
1635:
1628:
1626:
1620:
1618:
1615:
1608:
1607:
1604:
1600:
1594:
1590:
1584:
1580:
1578:
1574:
1572:
1568:
1566:
1562:
1560:
1556:
1554:
1550:
1549:
1546:
1541:
1533:
1528:
1526:
1521:
1519:
1514:
1513:
1510:
1502:
1498:
1496:
1490:
1487:
1484:
1481:
1476:
1472:
1471:
1458:
1454:
1449:
1444:
1440:
1436:
1432:
1425:
1423:
1421:
1419:
1417:
1407:
1402:
1398:
1394:
1390:
1388:
1379:
1371:
1367:
1363:
1359:
1354:
1349:
1345:
1341:
1337:
1330:
1322:
1318:
1314:
1308:
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962:
948:on 2011-03-11
947:
943:
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929:
918:
917:
910:
905:
902:
901:
894:
889:
886:
885:
878:
873:
870:
869:
868:Paraodontomma
862:
857:
854:
853:
852:Paraodontomma
846:
841:
838:
837:
830:
825:
822:
821:
814:
809:
806:
805:
798:
793:
792:
786:
781:
780:
769:
764:
763:
759:
753:
752:
751:Tetraphalerus
748:
742:
741:
737:
736:
733:Extant genera
727:
720:
719:
714:
710:Crowson, 1962
708:
707:
702:
696:
695:
690:
685:
680:
679:
674:
671:
664:
663:
658:
652:
651:
646:
641:
636:
635:
630:
624:
623:
622:Paraodontomma
618:
613:
608:
607:
602:
599:
592:
591:
586:
580:
579:
574:
568:
567:
562:
556:
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502:Jarzembowski
500:
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476:
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418:Jarzembowski
416:
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410:
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398:
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367:
363:
362:Burmese amber
356:
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316:
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304:
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263:
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261:Tetraphalerus
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114:
111:
110:Omma stanleyi
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
3439:
3318:Helophoridae
3310:Georissidae
3305:Epimetopidae
3221:Pleocomidae
3205:Ochodaeidae
3189:Hybosoridae
3173:Glaphyridae
3165:Geotrupidae
3143:
3140:Belohinidae
3101:Eucinetidae
3092:
3089:Decliniidae
3062:
3020:Phengodidae
2959:Cantharidae
2914:Dascillidae
2906:Dascilloidea
2888:Psephenidae
2880:Lutrochidae
2872:Limnichidae
2785:Buprestidae
2777:Buprestoidea
2717:Synchroidae
2701:Scraptiidae
2693:Salpingidae
2656:Prostomidae
2643:Oedemeridae
2619:Mordellidae
2537:Lymexylidae
2529:Lymexyloidea
2479:Attelabidae
2471:Anthribidae
2429:Smicripidae
2393:Phalacridae
2377:Nitidulidae
2372:Myraboliidae
2364:Monotomidae
2355:
2336:Kateretidae
2313:Cyclaxyridae
2273:
2270:Agapythidae
2234:Latridiidae
2184:Cerylonidae
2171:Anamorphidae
2105:Phycosecidae
2096:
2079:
2076:Metaxinidae
2029:Biphyllidae
2020:
1989:Orsodacnidae
1946:Cucujiformia
1879:Dermestidae
1749:
1688:Amphizoidae
1646:
1633:
1613:
1599:Archostemata
1583:Holometabola
1581:Superorder:
1575:Infraclass:
1500:
1494:
1488:Tree of Life
1438:
1434:
1396:
1392:
1386:
1378:
1343:
1339:
1329:
1301:
1294:
1261:
1257:
1247:
1194:
1190:
1179:
1144:
1140:
1119:the original
1114:
1105:
1064:
1060:
1050:
1025:
1021:
1015:
972:
968:
961:
950:. Retrieved
946:the original
932:
914:
898:
882:
866:
850:
834:
818:
804:Clessidromma
802:
777:
775:
767:
760:
749:
744:Newman, 1839
738:
725:
716:
704:
692:
683:
676:
669:
660:
648:
639:
632:
620:
611:
604:
597:
590:Notocupoides
588:
576:
564:
552:
540:
531:
524:
518:Zeuner, 1962
512:
503:
496:
484:
472:
465:
456:
449:
440:
428:
419:
414:Clessidromma
412:
400:
388:
376:
352:
342:Brochocoleus
340:
328:
319:
312:
302:Allophalerus
300:
289:
265:
259:
253:
238:Archostemata
229:
227:
209:
200:Archostemata
125:
109:
18:
3554:iNaturalist
3464:Wikispecies
3362:Hydraenidae
3336:Spercheidae
3323:Hydrochidae
3287:Synteliidae
3271:Histeridae
3263:Histeroidea
3213:Passalidae
3181:Glaresidae
3038:Throscidae
2998:(fireflies)
2996:Lampyridae
2983:Eucnemidae
2975:Elateridae
2935:Elateroidea
2751:Zopheridae
2635:Mycteridae
2566:Anthicidae
2437:Sphindidae
2421:Silvanidae
2331:Hobartiidae
2318:Erotylidae
2247:Mycetaeidae
2242:Murmidiidae
2226:Euxestidae
2120:Rentoniidae
1994:Oxypeltidae
1979:Disteniidae
1812:Lepiceridae
1741:Hygrobiidae
1733:Haliplidae
1717:Dytiscidae
1696:Aspidytidae
1141:Geosciences
998:11336/75307
694:Stegocoleus
662:Rhabdocupes
634:Pareuryomma
514:Liassocupes
458:Eurydictyon
430:Diluticupes
390:Cionocoleus
378:Burmocoleus
3641:Categories
3383:Silphidae
3375:Ptiliidae
3367:Leiodidae
3354:Agyrtidae
3197:Lucanidae
3076:Scirtoidea
3012:Omethidae
2991:Jurasaidae
2840:Dryopidae
2811:Byrrhidae
2803:Byrrhoidea
2495:Brentidae
2300:Cucujidae
2282:Boganiidae
2262:Cucujoidea
2135:Thymalidae
2125:Rhadalidae
2068:Melyridae
2037:Byturidae
1999:Vesperidae
1908:Ptiniidae
1758:Noteridae
1725:Gyrinidae
1701:Carabidae
1630:Jurodidae
1622:Cupedidae
1569:Subclass:
1559:Arthropoda
1540:Coleoptera
1321:1127945248
952:2009-11-21
924:References
836:Bukhkalius
779:Fuscicupes
678:Rhopalomma
566:Monticupes
542:Lithocupes
486:Kirejtomma
442:Echinocups
370:Cenomanian
354:Bukhkalius
346:Hong, 1982
196:Suborder:
190:Coleoptera
170:Arthropoda
3647:Ommatidae
3484:Ommatidae
3470:Ommatidae
3440:Ommatidae
3237:Trogidae
3231:(scarabs)
3109:Scirtidae
3084:Clambidae
2677:Pythidae
2611:Meloidae
2558:Aderidae
2461:(weevils)
2326:Helotidae
2252:Teredidae
2166:Alexiidae
2050:Cleridae
2009:Cleroidea
1842:Polyphaga
1840:Suborder
1793:Myxophaga
1791:Suborder
1746:Meruidae
1665:Suborder
1647:Ommatidae
1597:Suborder
1571:Pterygota
1551:Kingdom:
1486:Ommatidae
1480:Ommatidae
1457:2296-701X
1387:Notocupes
1370:264597717
1362:2624-2834
1221:0027-8424
1171:2076-3263
1147:(2): 73.
1097:250394641
1089:0375-0442
1028:: 34–39.
766:Escalona
762:Beutelius
726:Notocupes
718:Zygadenia
670:Notocupes
650:Polyakius
606:Odontomma
598:Notocupes
578:Notocupes
570:Ren, 1995
498:Lepidomma
466:Notocupes
450:Notocupes
434:Ren, 1995
402:Cionocups
394:Ren, 1995
272:Cupedidae
267:Beutelius
230:Ommatidae
210:Ommatidae
156:Kingdom:
150:Eukaryota
24:Ommatidae
3491:BioLib:
3449:Wikidata
3004:Lycidae
2848:Elmidae
2746:Ulodidae
2582:Boridae
2487:Belidae
2088:Peltidae
1681:families
1667:Adephaga
1577:Neoptera
1557:Phylum:
1553:Animalia
1542:families
1495:Ommatida
1286:84795808
1239:31740605
1007:21185393
884:Miniomma
820:Limnomma
554:Miniomma
526:Limnomma
330:Blapsium
246:Mesozoic
206:Family:
166:Phylum:
160:Animalia
146:Domain:
3533:2646448
3455:Q135044
2595:Ciidae
2503:Caridae
1565:Insecta
1563:Class:
1538:Extant
1266:Bibcode
1230:6900523
1199:Bibcode
1149:Bibcode
1069:Bibcode
1030:Bibcode
977:Bibcode
789:Gallery
682:Ashman
314:Asiania
234:beetles
186:Order:
180:Insecta
176:Class:
3611:112567
3598:110038
3585:678309
3572:109884
3559:467297
3507:200435
3494:165014
1679:Extant
1455:
1368:
1360:
1319:
1309:
1284:
1237:
1227:
1219:
1169:
1095:
1087:
1005:
770:, 2020
768:et al.
686:, 2015
684:et al.
642:, 2012
640:et al.
614:, 2006
612:et al.
534:, 2021
532:et al.
506:, 2019
504:et al.
422:, 2017
420:et al.
366:Albian
322:, 2023
320:et al.
286:Genera
250:Pangea
242:extant
221:, 1912
217:&
3624:69173
3567:IRMNG
1366:S2CID
1346:(5).
1282:S2CID
1093:S2CID
215:Sharp
3593:NCBI
3580:ITIS
3546:4750
3541:GBIF
3502:BOLD
1453:ISSN
1358:ISSN
1317:OCLC
1307:ISBN
1235:PMID
1217:ISSN
1167:ISSN
1085:ISSN
1003:PMID
740:Omma
638:Tan
610:Ren
318:Lee
264:and
255:Omma
228:The
219:Muir
35:PreꞒ
3528:EoL
3520:DL2
3515:CoL
3479:AFD
1443:doi
1401:doi
1348:doi
1274:doi
1225:PMC
1207:doi
1195:116
1157:doi
1077:doi
1065:323
1038:doi
993:hdl
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278:in
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70:T
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