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maximize the growth of its progeny, as the larger size represents greater competitiveness, so that the male lions could compete with other male lions. In order to control the size of the offspring within a certain range, the gene of the lioness will offset the growth-maximizing gene of the male lion. The genes of a female tiger, however, are not adapted to limiting growth, which allows ligers to grow extremely large—far larger and heavier than its parent species. In general, most ligers grow more than 3.3 m (10 ft 10 in) in length and weigh more than 400 kg (880 lb). According to the
365:
due to lack of a ruff or mane) with both stripes and "jaguar-like" rosettes on its sides. The assertion of hybrid identity was due to the combination of black, dark brown, light brown, dark orange, dark yellow and beige markings and the tiger-like stripes radiating from its face. It is more likely to have been a released liger, since these are very large and have a mix of rosettes (lion juvenile markings) and stripes and can have a brindled mix of colors exactly as described (their markings are extremely variable).
483:
25:
417:
closer set than those of an Indian leopard and were brown and indistinct, like the fading spots of a juvenile lion. The spots on the head, spine, belly and legs were black and distinct. The tail was spotted on the topside and striped underneath and had a blackish tip with longer hairs. The underside was dirty white, the ears were fawn and had a broad black bar, but did not have the white spot found in leopards.
374:
384:
292:
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hybrid, but most of them are just of strange-looking skins that could also be attributed to genetic mutations. Most of these reports are probably hoaxes or misinterpretations, which makes it hard for scientists to learn about tiger-leopardess hybrids, but at least a part of the claims are true or in part true, such as the ones made by
Frederick Codrington Hicks.
452:, which he translated as "bastard", and suggests a leopard/tiger hybrid (the reverse hybrid is unlikely to arise in the wild state, as a wild male tiger would probably kill rather than mate with a female leopard). Sankhala noted there was a belief amongst local people that leopards and tigers naturally hybridise.
810:
Guggisberg said liger and tigon were thought to be invariably sterile, which means they cannot have offspring. The first hybrid of a hybrid (that being a child of liger) was discovered at the Munich-Hellabrunn Zoo in 1943. The birth of the second generation of hybrids has proven that the biologists'
801:
A liger called Samson died at the age of thirteen in 2006. Shasta, a female liger, was born in the Hogle Zoo in Salt Lake City in 1948, and died in 1972. She lived for 24 years. Many claim that ligers are short-lived, but according to the survey, such a conclusion is still uncertain. A male tigon
792:
Ligers and tigons look just like their parents, only bigger or smaller. They have huge teeth, about two inches long. Their genes include the genetic components of tigers and lions, therefore, they may be very similar to tigers and lions, and can be difficult to identify. Their coloring ranges from
473:
In 1900, Carl
Hagenbeck crossed a female leopard with a Bengal tiger. The stillborn offspring had a mixture of spots, rosettes and stripes. Henry Scherren wrote, "A male tiger from Penang served two female Indian leopards, and twice with success. Details are not given and the story concludes somewhat
455:
From "The Tiger, Symbol Of
Freedom", edited by Nicholas Courtney: "Rare reports have been made of tigresses mating with leopards in the wild. There has even been an account of the sighting of rosettes; the stripes of the tiger being most prominent in the body. The animal was a male measuring a little
440:
Tiger-leopardess hybrids have supposedly appeared many times. Frederick
Codrington Hicks recorded that the weight of these creatures varied from 50 pounds to the weight of a tigress. In addition, in September 1965, a "leoger" skin was supposedly put on sale. There are some more documentations of this
360:
produced a male tiguar named Mickey. Mickey was on exhibition at a 400 m habitat and as of June 2009, was two years old and weighed 180 kg (400 lb). Attempts to verify this report have been bolstered by recent images purported to show the adult Mickey (see
External links section). There has
364:
There is a claimed sighting of a lion × black jaguar cross (male) and a tiger × black jaguar cross (female) loose in Maui, Hawaii. There are no authenticated tiger/jaguar hybrids and the description matches that of a liger. The alleged tiger × black jaguar was large, relatively long-necked (probably
723:
is the offspring of a female lion and a male tiger. The tigon is not as common as the converse hybrid, the liger. Contrary to some beliefs, the tigon ends up smaller than either parent, because male tigers and lionesses have a growth inhibitor. In the late 19th and early 20th centuries, tigons were
765:
Typically, the size of a liger is more likely to be larger and heavier than all other existing feline animals. Some biologists believe that the causes of its irregular large size, or 'gigantism', result from the lack of certain genes that limit the growth of lions. The male lion's genes tend to
416:
Natural
History Society. It was a cross between a large leopard and a lioness. Two cubs were born, one of which died aged 2.5 months, and the other was still living when Pocock described it in 1912. Pocock wrote that it was spotted like a leopard, but that the spots on its sides were smaller and
432:
is a native Indian name used for a supposedly natural hybrid offspring of a male leopard and a female tiger (tigress). Indian folklore claims that large male leopards sometimes mate with tigresses, and anecdotal evidence exists in India of offspring resulting from leopard to tigress matings. A
190:
In theory, lions and tigers can be matched in the wild and give offspring. In reality, there may be no natural born tigon or liger in the world, as both lions and tigers are separated both geographically and by behavioral differences. In
England, African lions and Asian tigresses have been
774:, US. He was measured at 3.33 m (10 ft 11 in), standing 1.25 m (4 ft 1 in) at the shoulder, and weighing 418.2 kg (922 lb). Hercules eats approximately 13.6 kg (30 lb) of meat per day, and drinks several liters of water per day.
242:
A leguar or lepjag is the hybrid of a male leopard and a female jaguar. The terms jagulep and lepjag are often used interchangeably, regardless of which animal was the sire. Numerous lepjags have been bred as animal actors, as they are more tractable than jaguars.
274:(Zoolog. Gart., 1863, 88). "The cub, a female, was grey. She is said to have produced two cubs to her sire; one like a jaguar, the other like the dam. Herr Rorig expressed his regret that the account of the last two cases mentioned lacked fullness and precision."
269:
in the panther (leopard) is recessive, the jaguar would have had to have been black, or be a jaguar-black leopard hybrid itself, carrying the recessive gene. Scherren continued, "The same cross, but with the sexes reversed, was noted, by
Professor Sacc (F) of
258:). These hybrids were also reared recently in Wombell's well-known travelling collection. I have seen some animals of this kind bred, between a male black jaguar and a female Indian leopard:-the young partook strongly of the male, being almost black."
819:
By 2017, roughly more than 100 ligers were thought to exist; however, only a few tigons still exist, as they are more difficult to breed. Moreover, ligers are more likely to attract tourists, so zoos prefer to breed ligers as opposed to tigons.
330:. Jahzara (female) and Tsunami (male) were the result of an unintended mating between a black jaguar called Diablo and a lioness called Lola, which had been hand-raised together and were inseparable. They were kept apart when Lola came into
210:
name, varying by which species is the sire (male parent) and which is the dam (female parent). For example, a hybrid between a lion and a tigress is a liger, because the lion is the male parent and the tigress is the female parent.
334:. Tsunami is spotted, but Jahzara is a melanistic jaglion due to inheriting the jaguar's dominant melanism gene. It was not previously known how the jaguar's dominant melanism gene would interact with lion coloration genes.
1444:
713:
is the offspring between a male lion and a female tiger, which is larger than its parents because the lion has a growth maximizing gene and the tigress, unlike the lioness, has no growth inhibiting gene.
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lamely. 'The leopardess dropped her cubs prematurely, the embryos were in the first stage of development and were scarcely as big as young mice.' Of the second leopardess there is no mention."
265:), a jaguar gave birth to two cubs from a union with a black leopard; one resembled the dam, but was somewhat darker, while the other was black with the rosettes of the dam showing. Since
191:
successfully mated, and three lion-tiger hybrid cubs were born in
Windsor in 1824, which is probably the earliest record of captive-bred ligers. The three cubs were then presented to
811:
knowledge of tigon and liger was wrong; it now seems that only male lion-tiger hybrids are invariably sterile; while female hybrids can give birth as other
Panthera animals as well.
702:(1961), Gerald Iles wrote "For the record I must say that I have never seen a liger, a hybrid obtained by crossing a lion with a tigress. They seem to be even rarer than tigons."
174:
research revealed that wild hybrids were also present in ancient times. The mitochondrial genomes of the snow leopard and the lion were more similar to each other than to other
1452:
781:, however inversely. They are smaller than the members of the parents species and weigh less than 150 kg (330 lb). A tigon is approximately twice as light as liger.
1044:
1279:(PhD). Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology. Uppsala: Acta Universitatis Upsaliensis. p. 8–10.
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Some zoos claim they breed ligers or tigons for conservation, but opponents believe that it is meaningless to preserve a species that does not exist in the wild.
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Unlike ligers, tigons are cross between a male tiger and a female lion, the presence of growth-minimizing genes from the lioness causing them to grow smaller.
281:. One such complex hybrid was exhibited in the early 1900s as a "Congolese spotted lion", hinting at some exotic African beast, rather than a man-made hybrid.
529:
Professor Valentine Bail conducted a long observation and recording of some lion-tiger hybrids, those lion-tiger are owned by Mr. Atkins and his zoo:
228:
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gold to brown to white, and they may have or not have spots or stripes. An adult male liger usually has a smaller mane than a male lion.
340:
When the fertile offspring of a male lion and female jaguar mates with a leopard, the resulting offspring is referred to as a leoliguar.
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1426:
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of male ancestors of modern snow leopards and female ancestors of modern lions interbred with male ancestors of modern snow leopards.
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1320:
Shankaranarayanan P, Banerjee M, Kacker RK, Aggarwal RK, Singh L (1997). "Genetic variation in Asiatic lions and Indian tigers".
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A jaglion or jaguon is the offspring between a male jaguar and a female lion (lioness). A mounted specimen is on display at the
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89:
61:
300:
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would not be perpetuated in the wild as the territories of the parental species do not overlap and the males are usually
348:
A tiguar is an offspring of a male tiger and a female jaguar. Reportedly, at the Altiplano Zoo in the city of San Pablo
1090:
770:(through 2013), the largest feline was the adult male liger, Hercules, from Myrtle Beach Safari, a wildlife reserve in
744:, mated with Debabrata, a male lion, and gave birth to three litigons. Only one litigon cub, named Cubanacan, survived.
68:
108:
466:
A tigard is the hybrid offspring of a tiger and a leopardess. The only known attempts to mate the two have produced
506:). The second generation hybrids of liger or tigon are known as liliger, tiliger, litigon and titigon. The tigon (
42:
75:
911:
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been no report of the birth of a hybrid from a male jaguar and female tiger, which would be termed a "jagger".
420:
Another lion-leopardess hybrid was born in Florence, Italy called lionard or lipard (/'laɪpəd/ or /'laɪpərd/).
277:
Female jaguleps or lepjags are fertile, and when one is mated to a male lion, the offspring are referred to as
46:
1985:
57:
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179:
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1149:
311:. It has the lion's background color, brown, jaguar-like rosettes and the powerful build of the jaguar.
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reported tiger/leopard matings were infertile, producing spontaneously aborted "walnut-sized fetuses".
2008:
1945:
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1980:
1950:
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35:
2013:
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947:: "In a paper on the breeding of the larger Felidae in captivity (P.Z.S. , 1861, p. 140),
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405:
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A jagupard, jagulep or jagleop is the hybrid of a male jaguar and a female leopard. A single
82:
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1500:
247:
171:
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A leopon is the result of breeding a leopard and a lioness. They occur only in captivity.
8:
1925:
864:"Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae)"
995:
888:
863:
734:
is the offspring of a lion and a ligress. The first known liliger is a cub named Kiara.
498:
The hybrids resulting from crossbreeding between lions and tigers are known as tigon (
352:(near Tlaxcala City, Mexico), the crossbreeding of a male Siberian tiger and a female
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893:
220:
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1975:
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163:
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Growth and Behaviour: Epigenetic and Genetic Factors Involved in Hybrid Dysgenesis
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1961:
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species, indicating that at some point in their history, the female hybrid
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1208:
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482:
1916:
1662:
1579:
741:
236:
207:
1045:"Cat Experts: Ligers and Other Designer Hybrids Pointless and Unethical"
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stated: "I have, more than once, met with instances of the male jaguar (
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1805:
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129:
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278:
192:
167:
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24:
1877:
1810:
1692:
1525:
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1513:
912:"Ligers and Tigons, Oh My! Cat Lineage Littered with Interbreeding"
842:
494:
Tiliger cross of (Panthera leo x Panthera tigris) x Panthera tigris
456:
over eight feet ." This is the same description as given by Hicks.
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266:
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A liguar is an offspring of a male lion and a female jaguar.
262:
143:
1755:
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owned by Atkins born on July 19, 1833, lived for 10 years.
383:
147:
1402:
756:
A titigon is the offspring of a male tiger and a tigoness.
291:
1564:
750:
A tiliger is the offspring of a male tiger and a ligress.
698:
The early record lion-tiger hybrid was mainly tigons, in
235:
were described as jagupards, which conforms to the usual
862:
Li, G.; Davis, B.W.; Eizirik, E.; Murphy, W.J. (2016).
1096:. International-Zoo-News. pp. 4–6. Archived from
861:
1119:
1117:
49:. Unsourced material may be challenged and removed.
2000:
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227:, United States. Jaguar-leopard hybrids bred at
855:
526:), as a hybrid of female tiger and male lion.
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974:
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214:
423:
314:On April 9, 2006, two jaglions were born at
295:Jaguar/lion hybrid, Rothschild Museum, Tring
1091:"Birth of a Lion × Leopard Hybrid in Italy"
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368:
343:
1492:
1478:
1403:"The Liger – Meet the World's Largest Cat"
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284:
887:
477:
448:refers to large, troublesome leopards as
223:female jagupard was produced at a zoo in
109:Learn how and when to remove this message
489:
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396:The first documented leopon was bred at
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132:between individuals of any of the five
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931:*H. Windischbauer, Hellbrun Zoo (1968)
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1125:"HYBRIDS BETWEEN LEOPARDS AND TIGERS"
1042:
1020:"HYBRIDS BETWEEN JAGUARS AND TIGERS"
522:). A liger is distinct from tigon (
198:
47:adding citations to reliable sources
18:
1267:
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1249:
1239:. London, Edward Lloyd. p. 45.
514:) is an offspring of a male tiger (
301:Walter Rothschild Zoological Museum
13:
1499:
1445:"The confusing world of the Liger"
1395:
1225:
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254:) breeding with a female leopard (
14:
2025:
1291:"Ligerungar – en världssensation"
1074:Pocock, R. I. (2 November 1912).
437:was reported in the early 1900s.
404:, in 1910. Its skin was sent to
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1313:
1297:. 3 August 2013. Archived from
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1043:Hall, Jani (24 February 2017).
34:needs additional citations for
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950:
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904:
814:
388:A group of taxidermied leopons
206:hybrids are typically given a
1:
1190:Guggisberg, C. A. W. (1975).
914:. Livescience.com. 2016-01-15
848:
787:
740:Rudrani, a tigoness from the
316:Bear Creek Wildlife Sanctuary
1355:"The litigon rediscovered".
1089:Florio, P. L. (March 1983).
805:
796:
7:
1270:"Hybrid dysgenesis effects"
826:
246:The 19th century zoologist
10:
2030:
1209:"liger | Size & Facts"
943:no. 2887, April 25, 1908,
261:In Barnabos Menagerie (in
215:Jaguar and leopard hybrids
185:
1986:Grizzly–polar bear hybrid
1981:Grizzly–black bear hybrid
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1934:
1914:
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1507:
1427:"Liger dies at sanctuary"
1177:no. 2887, April 25, 1908,
965:Baker, Taxonomy, pp. 5–7.
510:), also known as tiglon (
424:Leopard and tiger hybrids
1235:Sharpe, Bowdler (1897).
724:more common than ligers.
369:Leopard and lion hybrids
344:Jaguar and tiger hybrids
1334:10.1002/elps.1150180938
1213:Encyclopedia Britannica
1150:"Leopard-tiger hybrids"
777:Tigons too have growth
412:, the Secretary of the
285:Jaguar and lion hybrids
1846:Congolese spotted lion
1449:wildlifewaystation.org
1383:Guinness World Records
1365:10.1038/nindia.2017.46
1237:Lloyds Natural History
1192:Wild Cats of the World
975:Seymour, K.L. (1989).
956:Zoolog. Gart., 1861, 7
768:Guinness World Records
551:amount of female cubs
495:
487:
478:Lion and tiger hybrids
390:
380:
296:
1946:Polecat–ferret hybrid
880:10.1101/gr.186668.114
524:Panthera leo X tigris
518:) and a female lion (
508:Panthera tigris X leo
493:
485:
410:Walter Samuel Millard
406:Reginald Innes Pocock
386:
376:
294:
1379:"Largest living cat"
1250:Ile, Gerald (1961).
1055:on February 20, 2021
548:amount of male cubs
172:Mitochondrial genome
43:improve this article
1951:Polecat–mink hybrid
1431:Black Hills Pioneer
1049:National Geographic
533:
318:, Barrie (north of
239:naming convention.
1751:Lord Morton's mare
1252:At Home In The Zoo
700:At Home In The Zoo
608:December 31, 1826
532:
496:
488:
444:K Sankhala's book
391:
381:
356:from the southern
297:
16:Big cat crossbreed
1996:
1995:
1658:Jackal–dog hybrid
1570:Sheep–goat hybrid
1301:on August 4, 2013
984:Mammalian Species
696:
695:
562:October 24, 1824
461:Mammalian Hybrids
199:Naming of hybrids
119:
118:
111:
93:
58:"Panthera hybrid"
2021:
2009:Panthera hybrids
1976:ABC Islands bear
1906:Macropod hybrids
1883:Hybrid orangutan
1643:Dingo–dog hybrid
1494:
1487:
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1471:
1470:
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1451:. Archived from
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1328:(9): 1693–1700.
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1268:Shi, W. (2005).
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631:October 2, 1828
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378:Taxidermy leopon
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27:
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1826:Panthera hybrid
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1716:Elephant hybrid
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992:10.2307/3504096
979:
977:"Panthera onca"
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761:Growth and size
588:Clapham Common
585:April 22, 1825
545:number of cubs
542:place of birth
516:Panthera tigris
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371:
358:Lacandon Jungle
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1858:
1857:
1856:
1855:
1854:
1853:
1848:
1843:
1838:
1833:
1823:
1818:
1813:
1808:
1803:
1798:
1793:
1788:
1777:
1775:
1769:
1768:
1766:
1765:
1764:
1763:
1758:
1753:
1748:
1743:
1732:
1730:
1722:
1721:
1719:
1718:
1712:
1710:
1704:
1703:
1701:
1700:
1695:
1690:
1684:
1682:
1676:
1675:
1673:
1672:
1671:
1670:
1665:
1660:
1655:
1653:Eastern coyote
1650:
1645:
1640:
1635:
1624:
1622:
1616:
1615:
1613:
1612:
1607:
1602:
1596:
1594:
1588:
1587:
1585:
1584:
1583:
1582:
1577:
1572:
1567:
1562:
1557:
1552:
1550:American Breed
1541:
1539:
1533:
1532:
1529:
1528:
1522:
1516:
1509:
1508:
1505:
1504:
1501:Mammal hybrids
1497:
1496:
1489:
1482:
1474:
1466:
1465:
1436:
1418:
1407:ligerfacts.org
1394:
1370:
1347:
1312:
1295:Dagens Nyheter
1282:
1257:
1242:
1224:
1197:
1179:
1166:
1154:Macroevolution
1141:
1113:
1081:
1066:
1035:
1024:Messybeast.com
1011:
967:
958:
949:
945:Henry Scherren
933:
924:
903:
853:
852:
850:
847:
846:
845:
840:
835:
828:
825:
816:
813:
807:
804:
798:
795:
789:
786:
772:South Carolina
762:
759:
758:
757:
754:
751:
748:
745:
738:
735:
728:
725:
717:
714:
707:
694:
693:
690:
687:
684:
681:
678:
677:July 19, 1833
675:
671:
670:
667:
664:
661:
658:
655:
652:
648:
647:
644:
641:
638:
635:
632:
629:
628:fourth record
625:
624:
621:
618:
615:
612:
609:
606:
602:
601:
598:
595:
592:
589:
586:
583:
582:second record
579:
578:
575:
572:
569:
566:
563:
560:
556:
555:
552:
549:
546:
543:
540:
539:Date of birth
537:
479:
476:
459:The 1951 book
425:
422:
370:
367:
345:
342:
286:
283:
216:
213:
200:
197:
187:
184:
117:
116:
31:
29:
22:
15:
9:
6:
4:
3:
2:
2026:
2015:
2014:Felid hybrids
2012:
2010:
2007:
2006:
2004:
1987:
1984:
1982:
1979:
1977:
1974:
1973:
1972:
1969:
1968:
1966:
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1952:
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1927:
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1913:
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1844:
1842:
1839:
1837:
1834:
1832:
1829:
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1827:
1824:
1822:
1821:Serengeti cat
1819:
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1814:
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1809:
1807:
1804:
1802:
1799:
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1677:
1669:
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1659:
1656:
1654:
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1649:
1646:
1644:
1641:
1639:
1636:
1634:
1631:
1630:
1629:
1626:
1625:
1623:
1621:
1617:
1611:
1608:
1606:
1603:
1601:
1598:
1597:
1595:
1593:
1589:
1581:
1578:
1576:
1573:
1571:
1568:
1566:
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1551:
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1546:
1543:
1542:
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1538:
1534:
1527:
1523:
1521:
1517:
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1511:
1510:
1506:
1502:
1495:
1490:
1488:
1483:
1481:
1476:
1475:
1472:
1455:on 2019-03-01
1454:
1450:
1446:
1440:
1432:
1428:
1422:
1408:
1404:
1398:
1384:
1380:
1374:
1366:
1362:
1358:
1351:
1343:
1339:
1335:
1331:
1327:
1323:
1316:
1300:
1296:
1292:
1286:
1278:
1271:
1264:
1262:
1253:
1246:
1238:
1231:
1229:
1214:
1210:
1204:
1202:
1193:
1186:
1184:
1176:
1170:
1155:
1151:
1145:
1130:
1126:
1120:
1118:
1103:on 2014-10-22
1099:
1092:
1085:
1077:
1070:
1054:
1050:
1046:
1039:
1025:
1021:
1015:
1001:
997:
993:
989:
985:
978:
971:
962:
953:
946:
942:
937:
928:
913:
907:
899:
895:
890:
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881:
877:
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869:
865:
858:
854:
844:
841:
839:
836:
834:
831:
830:
824:
821:
812:
803:
794:
785:
782:
780:
775:
773:
769:
755:
752:
749:
746:
743:
739:
736:
733:
729:
726:
722:
718:
715:
712:
708:
705:
704:
703:
701:
691:
688:
685:
682:
679:
676:
674:sixth record
673:
672:
669:not recorded
668:
666:not recorded
665:
663:not recorded
662:
659:
656:
653:
651:fifth record
650:
649:
646:not recorded
645:
642:
639:
636:
633:
630:
627:
626:
623:a few months
622:
619:
616:
613:
610:
607:
605:third record
604:
603:
599:
597:not recorded
596:
594:not recorded
593:
590:
587:
584:
581:
580:
576:
573:
570:
567:
564:
561:
559:first record
558:
557:
553:
550:
547:
544:
541:
538:
536:
535:
530:
527:
525:
521:
517:
513:
509:
505:
502:) and liger (
501:
492:
484:
475:
471:
469:
464:
462:
457:
453:
451:
447:
442:
438:
436:
431:
421:
418:
415:
411:
407:
403:
399:
394:
389:
385:
379:
375:
366:
362:
359:
355:
351:
341:
338:
335:
333:
329:
325:
321:
317:
312:
310:
306:
305:Hertfordshire
302:
293:
289:
282:
280:
275:
273:
272:Barcelona Zoo
268:
264:
259:
257:
253:
249:
248:A.D. Bartlett
244:
240:
238:
234:
230:
226:
222:
212:
209:
205:
196:
194:
183:
181:
177:
173:
169:
165:
161:
157:
153:
149:
145:
141:
140:
136:of the genus
135:
131:
127:
125:
113:
110:
102:
99:February 2012
91:
88:
84:
81:
77:
74:
70:
67:
63:
60: –
59:
55:
54:Find sources:
48:
44:
38:
37:
32:This article
30:
26:
21:
20:
1971:Ursid hybrid
1960:
1935:
1926:Iron Age pig
1915:
1898:Macropodinae
1825:
1816:Savannah cat
1781:Felid hybrid
1736:Equid hybrid
1725:
1708:Elephantidae
1628:Canid hybrid
1545:Bovid hybrid
1457:. Retrieved
1453:the original
1448:
1439:
1430:
1421:
1410:. Retrieved
1406:
1397:
1386:. Retrieved
1382:
1373:
1357:Nature India
1356:
1350:
1325:
1321:
1315:
1303:. Retrieved
1299:the original
1294:
1285:
1276:
1251:
1245:
1236:
1216:. Retrieved
1212:
1191:
1174:
1169:
1157:. Retrieved
1153:
1144:
1132:. Retrieved
1128:
1105:. Retrieved
1098:the original
1084:
1075:
1069:
1057:. Retrieved
1053:the original
1048:
1038:
1027:. Retrieved
1023:
1014:
1003:. Retrieved
986:(340): 1–9.
983:
970:
961:
952:
940:
936:
927:
916:. Retrieved
906:
871:
867:
857:
833:Felid hybrid
822:
818:
809:
800:
791:
783:
776:
764:
699:
697:
528:
523:
520:Panthera leo
519:
515:
507:
497:
472:
465:
460:
458:
454:
449:
445:
443:
439:
434:
429:
427:
419:
395:
392:
387:
377:
363:
347:
339:
336:
313:
298:
288:
276:
260:
255:
251:
245:
241:
229:Hellbrun Zoo
218:
203:
202:
189:
175:
160:snow leopard
137:
123:
122:
120:
105:
96:
86:
79:
72:
65:
53:
41:Please help
36:verification
33:
1873:Denisova 11
1663:Sulimov dog
1159:January 15,
1134:October 16,
874:(1): 1–11.
815:Zoo animals
742:Alipore Zoo
657:Kensington
600:short time
450:adhabaghera
350:Apetatitlán
237:portmanteau
208:portmanteau
2003:Categories
1888:Koolakamba
1806:Kellas cat
1786:Bengal cat
1560:Dwarf Lulu
1459:2019-05-17
1412:2019-05-13
1388:2019-05-13
1218:2019-05-13
1129:Messybeast
1107:2015-01-11
1029:2016-04-06
1005:2015-09-09
918:2016-04-06
849:References
788:Appearance
680:Liverpool
654:May, 1831
611:Edinburgh
554:longevity
512:/ˈtaɪɡlən/
468:stillborns
279:lijaguleps
130:crossbreed
69:newspapers
1865:Hominidae
1592:Camelidae
1175:The Field
1076:The Field
941:The Field
806:Fertility
797:Longevity
779:dysplasia
692:10 years
504:/ˈlaɪɡər/
500:/ˈtaɪɡən/
433:supposed
428:The name
256:P. pardus
193:George IV
168:infertile
1878:Humanzee
1811:Pumapard
1693:Wholphin
1526:Mammalia
1520:Chordata
1514:Animalia
1512:Kingdom
1359:. 2017.
1305:4 August
1059:29 April
898:26518481
843:Pumapard
827:See also
634:Windsor
565:Windsor
398:Kolhapur
267:melanism
233:Salzburg
221:rosetted
204:Panthera
176:Panthera
162:. Most
139:Panthera
124:Panthera
1937:Mustela
1836:Liliger
1801:Chausie
1796:Caraval
1773:Felidae
1761:Zebroid
1698:Narluga
1680:Cetacea
1668:Wolfdog
1638:Coywolf
1620:Canidae
1610:Huarizo
1555:Beefalo
1537:Bovidae
1518:Phylum
1342:9378147
1000:3504096
889:4691742
753:Titigon
747:Tiliger
737:Litigon
732:liliger
727:Liliger
577:1 year
332:oestrus
324:Ontario
320:Toronto
309:England
252:P. onca
225:Chicago
186:History
180:progeny
164:hybrids
156:leopard
134:species
83:scholar
1841:Marozi
1648:Dogxim
1633:Coydog
1580:Żubroń
1575:Yakalo
1524:Class
1340:
998:
896:
886:
838:Marozi
414:Bombay
354:jaguar
328:Canada
158:, and
152:jaguar
142:: the
126:hybrid
85:
78:
71:
64:
56:
1962:Ursus
1851:Tigon
1831:Liger
1791:Blynx
1746:Kunga
1741:Hinny
1727:Equus
1605:Bukht
1273:(PDF)
1101:(PDF)
1094:(PDF)
996:JSTOR
980:(PDF)
721:tigon
716:Tigon
711:liger
706:Liger
486:Liger
446:Tiger
435:dogla
430:dogla
402:India
263:Spain
144:tiger
128:is a
90:JSTOR
76:books
1756:Mule
1600:Cama
1338:PMID
1307:2013
1161:2015
1136:2023
1061:2023
894:PMID
148:lion
62:news
1917:Sus
1565:Dzo
1361:doi
1330:doi
1173:In
988:doi
884:PMC
876:doi
408:by
322:),
45:by
2005::
1447:.
1429:.
1405:.
1381:.
1336:.
1326:18
1324:.
1293:.
1275:.
1260:^
1227:^
1211:.
1200:^
1182:^
1152:.
1127:.
1116:^
1047:.
1022:.
994:.
982:.
892:.
882:.
872:26
870:.
866:.
730:A
719:A
709:A
689:2
686:1
683:3
660:3
643:2
640:1
637:3
620:2
617:1
614:3
591:3
574:1
571:2
568:3
470:.
400:,
326:,
307:,
303:,
231:,
195:.
170:.
154:,
150:,
146:,
121:A
1493:e
1486:t
1479:v
1462:.
1433:.
1415:.
1391:.
1367:.
1363::
1344:.
1332::
1309:.
1254:.
1221:.
1194:.
1163:.
1138:.
1110:.
1078:.
1063:.
1032:.
1008:.
990::
921:.
900:.
878::
112:)
106:(
101:)
97:(
87:·
80:·
73:·
66:·
39:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.